In order to locate QTLs controlling the phenotypic stability and drought tolerance of yield and yield components in barley, seven disomic addition lines were sown together with their parents (donor and recipient) in a randomized complete block design with three replications under four rainfed and irrigated conditions. The descriptive diagram of yield and yield components exhibited a genotype (G) × environment (E) interaction and moderate variability over different environments, indicating the possibility of selection for stable and drought-tolerant entries. The AMMI stability value (ASV) and yield stability index (YSI) discriminated addition lines 2H and 4H as the most stable and droughttolerant.Path analysis revealed that the relative contribution of the number of seeds per plant (NSPP) (0.71) to grain yield (GY) was higher than that of the number of seeds per spike (SPS) (−0.44) and of thousand-seed weight (TSW) (−0.14). Therefore, the contribution of NSPP to the stability of GY over different environments was higher than that of other yield components. In other words, the instability of GY was caused by TSW and SPS in different environments. Path analysis on the drought susceptibility index revealed that most of the QTLs controlling drought tolerance and drought susceptibility in barley are located on chromosomes 3H and 6H, respectively.
Askarinia P., Saeidi G., Rezaei A. (2008): Assessment of genotype × environment interaction in ten wheat cultivars with regression and path coefficient analysis. Electronic J. Crop Prod., 1, 64–81.
Rezaei A. , 'Assessment of genotype × environment interaction in ten wheat cultivars with regression and path coefficient analysis ' (2008 ) 1 Electronic J. Crop Prod. : 64 -81 .
Atta B. M., Shah T. M. (2009): Stability analysis of elite chickpea genotypes tested under diverse environments. Aust. J. Crop Sci., 3, 249–256.
Shah T. M. , 'Stability analysis of elite chickpea genotypes tested under diverse environments ' (2009 ) 3 Aust. J. Crop Sci. : 249 -256 .
Babarmanzoor A., Tariq M. S., Ghulam A., Muhammad A. (2009): Genotype × environment interaction for seed yield in Kabuli Chickpea (Cicer arietinum L.) genotypes. Appl. Genet.., 96, 397–405.
Muhammad A. , 'Genotype × environment interaction for seed yield in Kabuli Chickpea (Cicer arietinum L.) genotypes ' (2009 ) 96 Appl. Genet.. : 397 -405 .
Bajpai P. K., Prabhakaran V. T. (2000): A new procedure of simultaneous selection for high yielding and stable crop genotypes. Indian J. Genet., 60, 141–146.
Prabhakaran V. T. , 'A new procedure of simultaneous selection for high yielding and stable crop genotypes ' (2000 ) 60 Indian J. Genet. : 141 -146 .
Bennett M. D., Smith L. B. (1976): Nuclear DNA amounts in angiosperms. Philosophical Transactions of the Royal Society (London). Biol. Sci., 274, 227–274.
Smith L. B. , 'Nuclear DNA amounts in angiosperms ' (1976 ) 274 Philosophical Transactions of the Royal Society (London). Biol. Sci. : 227 -274 .
Busch R. H., Hammond J., Frohberg R. C. (1976): Stability and performance of hard red spring wheat bulks for grain yield. Crop Sci., 16, 256–259.
Frohberg R. C. , 'Stability and performance of hard red spring wheat bulks for grain yield ' (1976 ) 16 Crop Sci. : 256 -259 .
Darvishzadeh R., Hatami-Maleki H., Sarrafi A. (2011): Path analysis of the relationships between yield and some related traits in diallel population of sunflower (Helianthus annuus L.) under well-watered and water-stressed conditions. Austral. J. Crop Sci., 5, 674–680.
Sarrafi A. , 'Path analysis of the relationships between yield and some related traits in diallel population of sunflower (Helianthus annuus L.) under well-watered and water-stressed conditions ' (2011 ) 5 Austral. J. Crop Sci. : 674 -680 .
Das M. K., Taliaferro C. M. (2009): Genetic variability and interrelationships of seed yield and yield components in switchgrass. Euphytica, 167, 95–105.
Taliaferro C. M. , 'Genetic variability and interrelationships of seed yield and yield components in switchgrass ' (2009 ) 167 Euphytica : 95 -105 .
Dhillon B. S., Singh J. (1977): Estimation and inheritance of stability parameters of grain yield in maize. J. Agric. Sci., 88, 256–257.
Singh J. , 'Estimation and inheritance of stability parameters of grain yield in maize ' (1977 ) 88 J. Agric. Sci. : 256 -257 .
Ebadi Segherloo A., Sabaghpour S. H., Dehghani H., Kamrani M. (2008): Non-parametric measures of phenotypic stability in chickpea genotypes (Cicer arietinum L.). Euphytica, 162, 221–229.
Kamrani M. , 'Non-parametric measures of phenotypic stability in chickpea genotypes (Cicer arietinum L.) ' (2008 ) 162 Euphytica : 221 -229 .
Ellis R. P., Forster B. P., Robinson D., Handly L. L., Gordon D. C. (2000): Wild barley: A source of genes for crop improvement in the 21st century. J. Exp. Bot., 51, 9–17.
Gordon D. C. , 'Wild barley: A source of genes for crop improvement in the 21st century ' (2000 ) 51 J. Exp. Bot. : 9 -17 .
Eskridge K. M., Shah M. M., Baenziger P. S., Travnicek D. A. (2000): Correcting for classification errors when estimating the number of genes using recombinant inbred chromosome lines. Crop Sci., 40, 398–403.
Travnicek D. A. , 'Correcting for classification errors when estimating the number of genes using recombinant inbred chromosome lines ' (2000 ) 40 Crop Sci. : 398 -403 .
Fagam A. S., Bununu A. M., Buba U. M. (2006): Path coefficient analysis of the components of grain yield in wheat (Triticum aestivum L.). Int. J. Nat. Appl. Sci., 2, 336–340.
Buba U. M. , 'Path coefficient analysis of the components of grain yield in wheat (Triticum aestivum L.) ' (2006 ) 2 Int. J. Nat. Appl. Sci. : 336 -340 .
Farshadfar E. (2008): Incorporation of AMMI stability value and grain yield in a single nonparametric index (GSI) in bread wheat. Pak. J. Biol. Sci., 11, 1791–1796.
Farshadfar E. , 'Incorporation of AMMI stability value and grain yield in a single nonparametric index (GSI) in bread wheat ' (2008 ) 11 Pak. J. Biol. Sci. : 1791 -1796 .
Farshadfar E. (2013): Simultaneous selection of yield and yield stability in chickpea genotypes using the GGE biplot technique. Acta Agron. Hung., 61, 185–194.
Farshadfar E. , 'Simultaneous selection of yield and yield stability in chickpea genotypes using the GGE biplot technique ' (2013 ) 61 Acta Agron. Hung. : 185 -194 .
Farshadfar E., Farshadfar M., Sutka J. (1999): Genetic analysis of phenotypic stability parameters in wheat. Acta Agron. Hung., 47, 109–116.
Sutka J. , 'Genetic analysis of phenotypic stability parameters in wheat ' (1999 ) 47 Acta Agron. Hung. : 109 -116 .
Farshadfar E., Farshadfar M., Sutka J. (2000): Combining ability analysis of drought tolerance in wheat over different water regimes. Acta Agron. Hung., 48, 353–361.
Sutka J. , 'Combining ability analysis of drought tolerance in wheat over different water regimes ' (2000 ) 48 Acta Agron. Hung. : 353 -361 .
Farshadfar E., Mohammad Mehdi P., Maryam J. (2012): Evaluation of phenotypic stability in bread wheat genotypes using GGE-biplot. Int. J. Agric. Crop Sci., 4, 904–910.
Maryam J. , 'Evaluation of phenotypic stability in bread wheat genotypes using GGE-biplot ' (2012 ) 4 Int. J. Agric. Crop Sci. : 904 -910 .
Farshadfar E., Sutka J. (2003): Locating QTLs controlling adaptation in wheat using AMMI model. Cereal Res. Commun., 31, 249–254.
Sutka J. , 'Locating QTLs controlling adaptation in wheat using AMMI model ' (2003 ) 31 Cereal Res. Commun. : 249 -254 .
Gauch H. G., Zobel R. W. (1996): AMMI analysis of yield trials. In: Kang M. S., Gauch H. G. (eds.), Genotype by Environment Interaction. CRC Press, Boca Raton, FL.
Zobel R. W. , '', in Genotype by Environment Interaction , (1996 ) -.
Grafius J. E., Thomas R. L. (1971): The case for indirect genetic control of sequential traits and the strategy of deployment of environmental resources of the plant. Heredity, 26, 433–442.
Thomas R. L. , 'The case for indirect genetic control of sequential traits and the strategy of deployment of environmental resources of the plant ' (1971 ) 26 Heredity : 433 -442 .
Hui Z., Bin Z. Z., Shao H. B., Xu P., Foulkes M. J. (2008): Genetic correlation and path analysis of transpiration efficiency for wheat flag leaves. Envir. Exper. Bot., 64, 128–134.
Foulkes M. J. , 'Genetic correlation and path analysis of transpiration efficiency for wheat flag leaves ' (2008 ) 64 Envir. Exper. Bot. : 128 -134 .
Jalata Z. (2011): GGE-biplot analysis of multi-environment yield trials of barley (Hordeum vulgare L.) genotypes in Southeastern Ethiopia. Inter. J. Plant Breed. Genet., 5, 59–75.
Jalata Z. , 'GGE-biplot analysis of multi-environment yield trials of barley (Hordeum vulgare L.) genotypes in Southeastern Ethiopia ' (2011 ) 5 Inter. J. Plant Breed. Genet. : 59 -75 .
Kang M. S. (1993): Simultaneous selection for yield and stability in crop performance trials: Consequences for growers. Agron. J., 85, 754–757.
Kang M. S. , 'Simultaneous selection for yield and stability in crop performance trials: Consequences for growers ' (1993 ) 85 Agron. J. : 754 -757 .
Khazaie H., Mohammady S., Monneveux P., Stoddard F. (2011): The determination of direct and indirect effects of carbon isotope discrimination (Δ), stomatal characteristics and water use efficiency on grain yield in wheat using sequential path analysis. Aust. J. Crop Sci., 5, 466–472.
Stoddard F. , 'The determination of direct and indirect effects of carbon isotope discrimination (Δ), stomatal characteristics and water use efficiency on grain yield in wheat using sequential path analysis ' (2011 ) 5 Aust. J. Crop Sci. : 466 -472 .
Kőszegi B., Farshadfar E., Vágújfalvi A., Sutka J. (1996): Drought tolerance studies on wheat/rye disomic chromosome addition lines. Acta Agron. Hung., 44, 121–126.
Sutka J. , 'Drought tolerance studies on wheat/rye disomic chromosome addition lines ' (1996 ) 44 Acta Agron. Hung. : 121 -126 .
Lin C. S., Binns M. R. (1988): A method of analyzing cultivar × location × year experiments: a new stability parameter. Theor. Appl. Genet., 76, 425–430.
Binns M. R. , 'A method of analyzing cultivar × location × year experiments: a new stability parameter ' (1988 ) 76 Theor. Appl. Genet. : 425 -430 .
Lin C. S., Binns M. R. (1991): Genetic properties of four stability parameters. Theor. Appl. Genet., 82, 505–509.
Binns M. R. , 'Genetic properties of four stability parameters ' (1991 ) 82 Theor. Appl. Genet. : 505 -509 .
Lin C. S., Binns M. R. (1994): Concepts and methods for analyzing regional trial data for cultivar and location selection. Plant Breed. Rev., 12, 271–297.
Binns M. R. , 'Concepts and methods for analyzing regional trial data for cultivar and location selection ' (1994 ) 12 Plant Breed. Rev. : 271 -297 .
Lynch D. R., Tai G. C. C. (1989): Yield and yield component response of eight potato genotypes to water stress. Crop Sci., 29, 1207–1211.
Tai G. C. C. , 'Yield and yield component response of eight potato genotypes to water stress ' (1989 ) 29 Crop Sci. : 1207 -1211 .
Mohammadi R., Abdulahi A., Haghparast R., Armion M. (2007): Interpreting genotypeenvironment interactions for durum wheat grain yields using non-parametric methods. Euphytica, 157, 239–251.
Armion M. , 'Interpreting genotypeenvironment interactions for durum wheat grain yields using non-parametric methods ' (2007 ) 157 Euphytica : 239 -251 .
Mohammadi R., Amri A. (2008): Comparison of parametric and non-parametric methods for selecting stable and adapted durum wheat genotypes in variable environments. Euphytica, 159, 419–432.
Amri A. , 'Comparison of parametric and non-parametric methods for selecting stable and adapted durum wheat genotypes in variable environments ' (2008 ) 159 Euphytica : 419 -432 .
Mohammadinejad G., Rezaei A. (2006): Genotype and environment interaction (agronomy managements) in oat cultivars on the base of path and regression analysis. J. Sci. Tech. Agric. Natur. Res., 11, 187–198.
Rezaei A. , 'Genotype and environment interaction (agronomy managements) in oat cultivars on the base of path and regression analysis ' (2006 ) 11 J. Sci. Tech. Agric. Natur. Res. : 187 -198 .
Morgan M. (1991): A gene controlling difference in osmoregulation in wheat. Aust. J. Plant Physiol., 18, 249–257.
Morgan M. , 'A gene controlling difference in osmoregulation in wheat ' (1991 ) 18 Aust.^J. Plant Physiol. : 249 -257 .
Patanothai A., Atkins R. E. (1974): Yield stability of single crosses and three-way hybrids of grain sorghum. Crop Sci., 14, 288–290.
Atkins R. E. , 'Yield stability of single crosses and three-way hybrids of grain sorghum ' (1974 ) 14 Crop Sci. : 288 -290 .
Purchase J. L., Hatting H., Van Deventer C. S. (2000): Genotype × environment interaction of winter wheat in South Africa: II. Stability analysis of yield performance. South Afric. J. Plant Soil., 17, 101–107.
Deventer C. S. , 'Genotype × environment interaction of winter wheat in South Africa: II. Stability analysis of yield performance ' (2000 ) 17 South Afric. J. Plant Soil. : 101 -107 .
Rao A. R., Prabhakaran V. T. (2005): Use of AMMI in simultaneous selection of genotypes for yield and stability. Ind. Soc. Agril. Statist., 59, 76–82.
Prabhakaran V. T. , 'Use of AMMI in simultaneous selection of genotypes for yield and stability ' (2005 ) 59 Ind. Soc. Agril. Statist. : 76 -82 .
Shpiler L., Blum A. (1991): Heat tolerance for yield and its components in different wheat cultivars. Euphytica, 51, 257–263.
Blum A. , 'Heat tolerance for yield and its components in different wheat cultivars ' (1991 ) 51 Euphytica : 257 -263 .
Tai G. C. C. (1975): Analysis of genotype environment interactions based on the method of path coefficient analysis. Can. J. Genet. Cytol., 17, 141–149.
Tai G. C. C. , 'Analysis of genotype environment interactions based on the method of path coefficient analysis ' (1975 ) 17 Can. J. Genet. Cytol. : 141 -149 .
Tai G. C. C. (1979): Analysis of genotype environment interaction of potato yield. Crop Sci., 19, 434–438.
Tai G. C. C. , 'Analysis of genotype environment interaction of potato yield ' (1979 ) 19 Crop Sci. : 434 -438 .
Tai G. C. C. (1990): Path analysis of genotype-environment interactions. pp. 273–286. In: Kang M. S. (ed.), Genotype-By-Environment Interaction and Plant Breeding. Louisiana University, Baton Rouge.
Tai G. C. C. , '', in Genotype-By-Environment Interaction and Plant Breeding , (1990 ) -.
Tai G. C. C., Levy D., Coleman W. K. (1994): Path analysis of genotype-environment interaction of potatoes exposed to increasingly warm climate constraints. Euphytica, 75, 49–61.
Coleman W. K. , 'Path analysis of genotype-environment interaction of potatoes exposed to increasingly warm climate constraints ' (1994 ) 75 Euphytica : 49 -61 .
Yaghotipoor A., Farshadfar E. (2007): Non-parametric estimation and component analysis of phenotypic stability in chickpea (Cicer arietinum L.). Pak. J. Biol. Sci., 10, 2646–2652.
Farshadfar E. , 'Non-parametric estimation and component analysis of phenotypic stability in chickpea (Cicer arietinum L.) ' (2007 ) 10 Pak. J. Biol. Sci. : 2646 -2652 .
Yasin A. B., Singh S. (2010): Correlation and path coefficient analyses in sunflower. J. Plant Breed. Crop Sci., 2, 129–133.
Singh S. , 'Correlation and path coefficient analyses in sunflower ' (2010 ) 2 J. Plant Breed. Crop Sci. : 129 -133 .
Zali, H., Farshadfar, E., Sabaghpour, S. H. (2011): Nonparametric analysis of phenotypic stability in chickpea (Cicer arietinum L.) genotypes in Iran. Crop Breed. J., 1, 89–100.
Sabaghpour S. H. , 'Nonparametric analysis of phenotypic stability in chickpea (Cicer arietinum L.) ' (2011 ) 1 Crop Breed. J. : 89 -100 .
Zobel R. W., Wright M. J., Gauch H. G. (1988): Statistical analysis of a yield trial. Agron. J., 80, 388–393.
Gauch H. G. , 'Statistical analysis of a yield trial ' (1988 ) 80 Agron.^J. : 388 -393 .