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  • 1 School of Psychology, University of Birmingham, Edgbaston, Birmingham, B15 2TT, UK
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Abstract

In our everyday interactions we encounter a plethora of novel experiences in different contexts that require prompt decisions for successful actions and social interactions. Despite the seeming ease with which we perform these interactions, extracting the key information from the highly complex input of the natural world and deciding how to interpret it is a computationally demanding task for the visual system. Accumulating evidence suggests that the brain solves this problem by combining sensory information and previous knowledge about the environment. Here, we review the neural mechanisms that mediate experience-based plasticity and shape perceptual decisions. We propose that learning plays an important role in the adaptive optimization of visual functions that translate sensory experiences to decisions by shaping neural representations across cortical circuits in the primate brain.

  • M. Ahissar S. Hochstein 2004 The reverse hierarchy theory of visual perceptual learning Trends Cogn. Sci. 8 10 457 464.

  • F. G. Ashby W. T. Maddox 2005 Human category learning Annu. Rev. Psychol. 56 149.

  • C. I. Baker M. Behrmann C. R. Olson 2002 Impact of learning on representation of parts and wholes in monkey inferotemporal cortex Nat. Neurosci. 5 11 1210 1216.

    • Search Google Scholar
    • Export Citation
  • H. Barlow 1990 Conditions for versatile learning, Helmholtz's unconscious inference, and the task of perception Vision Res. 30 11 1561 1571.

    • Search Google Scholar
    • Export Citation
  • M. C. A. Booth E. T. Rolls 1998 View-invariant representations of familiar objects by neurons in the inferior temporal visual cortex Cerebral Cortex 8 510 523.

    • Search Google Scholar
    • Export Citation
  • W. H. Bosking Y. Zhang B. Schofield D. Fitzpatrick 1997 Orientation selectivity and the arrangement of horizontal connections in tree shrew striate cortex J. Neurosci. 17 6 2112.

    • Search Google Scholar
    • Export Citation
  • M. J. Brady D. Kersten 2003 Bootstrapped learning of novel objects J. Vis. 3 6 413 422.

  • C. Buchel J. T. Coull K. J. Friston 1999 The predictive value of changes in effective connectivity for human learning Science 283 5407 1538 1541.

    • Search Google Scholar
    • Export Citation
  • L. L. Chao J. Weisberg A. Martin 2002 Experience-dependent modulation of category-related cortical activity Cereb Cortex 12 5 545 551.

  • H. J. Chisum D. Fitzpatrick 2004 The contribution of vertical and horizontal connections to the receptive field center and surround in V1 Neural Netw. 17 5–6 681.

    • Search Google Scholar
    • Export Citation
  • M. Corbetta G. L. Shulman 2002 Control of goal-directed and stimulus-driven attention in the brain Nat. Rev. Neurosci. 3 3 201 215.

  • D. D. Cox R. L. Savoy 2003 Functional magnetic resonance imaging (fMRI) “brain reading”: Detecting and classifying distributed patterns of fMRI activity in human visual cortex Neuroimage 19 2Pt1 261.

    • Search Google Scholar
    • Export Citation
  • R. E. Crist W. Li C. D. Gilbert 2001 Learning to see: Experience and attention in primary visual cortex Nat. Neurosci. 4 5 519 525.

  • R. Desimone 1998 Visual attention mediated by biased competition in extrastriate visual cortex Philos. Trans. R. Soc. Lond. B Biol. Sci. 353 1373 1245 1255.

    • Search Google Scholar
    • Export Citation
  • R. Desimone J. Duncan 1995 Neural mechanisms of selective visual attention Annu. Rev. Neurosci. 18 193 222.

  • R. J. Dolan G. R. Fink E. Rolls M. Booth A. Holmes R. S. Frackowiak K. J. Friston 1997 How the brain learns to see objects and faces in an impoverished context Nature 389 6651 596 599.

    • Search Google Scholar
    • Export Citation
  • B. A. Dosher Z. L. Lu 1998 Perceptual learning reflects external noise filtering and internal noise reduction through channel reweighting Proc. Natl. Acad. Sci. USA 95 23 13988 13993.

    • Search Google Scholar
    • Export Citation
  • J. Duncan 2001 An adaptive coding model of neural function in prefrontal cortex Nat. Rev. Neurosci. 2 11 820 829.

  • J. Duncan A. M. Owen 2000 Common regions of the human frontal lobe recruited by diverse cognitive demands Trends Neurosci. 23 10 475 483.

    • Search Google Scholar
    • Export Citation
  • M. P. Eckstein C. K. Abbey B. T. Pham S. S. Shimozaki 2004 Perceptual learning through optimization of attentional weighting: Human versus optimal Bayesian learner J. Vis. 4 12 1006 1019.

    • Search Google Scholar
    • Export Citation
  • M. Fahle 2004 Perceptual learning: A case for early selection J. Vis. 4 10 879 890.

  • I. Fine R. A. Jacobs 2002 Comparing perceptual learning tasks: Areview J. Vis. 2 2 190 203.

  • D. Fitzpatrick 2000 Seeing beyond the receptive field in primary visual cortex Curr. Opin. Neurobiol. 10 4 438 443.

  • D. J. Freedman M. Riesenhuber T. Poggio E. K. Miller 2003 A comparison of primate prefrontal and inferior temporal cortices during visual categorization J. Neurosci. 23 12 5235 5246.

    • Search Google Scholar
    • Export Citation
  • C. S. Furmanski D. Schluppeck S. A. Engel 2004 Learning strengthens the response of primary visual cortex to simple patterns Curr. Biol. 14 7 573 578.

    • Search Google Scholar
    • Export Citation
  • I. Gauthier A. W. Anderson M. J. Tarr P. Skudlarski J. C. Gore 1997 Levels of categorization in visual recognition studied using functional magnetic resonance imaging Curr. Biol. 7 9 645 651.

    • Search Google Scholar
    • Export Citation
  • I. Gauthier M. J. Tarr A. W. Anderson P. Skudlarski J. C. Gore 1999 Activation of the middle fusiform ‘face area’ increases with expertise in recognizing novel objects Nat. Neurosci. 2 6 568 573.

    • Search Google Scholar
    • Export Citation
  • W. S. Geisler J. S. Perry B. J. Super D. P. Gallogly 2001 Edge co-occurrence in natural images predicts contour grouping performance Vision Res. 41 6 711 724.

    • Search Google Scholar
    • Export Citation
  • G. M. Ghose T. Yang J. H. Maunsell 2002 Physiological correlates of perceptual learning in monkey V1 and V2 J. Neurophysiol. 87 4 1867 1888.

    • Search Google Scholar
    • Export Citation
  • C. D. Gilbert 1992 Horizontal integration and cortical dynamics Neuron 9 1 1 13.

  • C. D. Gilbert M. Sigman R. E. Crist 2001 The neural basis of perceptual learning Neuron 31 5 681 697.

  • C. D. Gilbert T. N. Wiesel 1989 Columnar specificity of intrinsic horizontal and corticocortical connections in cat visual cortex J. Neurosci. 9 7 2432.

    • Search Google Scholar
    • Export Citation
  • P. W. Glimcher 2003 The neurobiology of visual-saccadic decision making Annu. Rev. Neurosci. 26 133 179.

  • J. Gold P. J. Bennett A. B. Sekuler 1999 Signal but not noise changes with perceptual learning Nature 402 6758 176 178.

  • J. I. Gold M. N. Shadlen 2003 The influence of behavioral context on the representation of a perceptual decision in developing oculomotor commands J. Neurosci. 23 2 632 651.

    • Search Google Scholar
    • Export Citation
  • J. I. Gold M. N. Shadlen 2007 The neural basis of decision making Annu. Rev. Neurosci. 30 535 574.

  • R. L. Goldstone 1998 Perceptual learning Annu. Rev. Psychol. 49 585 612.

  • R. L. Goldstone Y. Lippa R. M. Shiffrin 2001 Altering object representations through category learning Cognition 78 1 27 43.

  • K. Grill-Spector T. Kushnir T. Hendler R. Malach 2000 The dynamics of object-selective activation correlate with recognition performance in humans Nat. Neurosci. 3 8 837 843.

    • Search Google Scholar
    • Export Citation
  • J. Grinband J. Hirsch V. P. Ferrera 2006 A neural representation of categorization uncertainty in the human brain Neuron 49 5 757 763.

  • S. J. Hanson T. Matsuka J. V. Haxby 2004 Combinatorial codes in ventral temporal lobe for object recognition: Haxby (2001) revisited: Is there a “face” area? Neuroimage 23 1 156.

    • Search Google Scholar
    • Export Citation
  • J. V. Haxby M. I. Gobbini M. L. Furey A. Ishai J. L. Schouten P. Pietrini 2001 Distributed and overlapping representations of faces and objects in ventral temporal cortex Science 293 5539 2425 2430.

    • Search Google Scholar
    • Export Citation
  • J. D. Haynes G. Rees 2006 Decoding mental states from brain activity in humans Nat. Rev. Neurosci. 7 7 523.

  • H. R. Heekeren S. Marrett P. A. Bandettini L. G. Ungerleider 2004 A general mechanism for perceptual decision-making in the human brain Nature 431 7010 859 862.

    • Search Google Scholar
    • Export Citation
  • H. R. Heekeren S. Marrett D. A. Ruff P. A. Bandettini L. G. Ungerleider 2006 Involvement of human left dorsolateral prefrontal cortex in perceptual decision making is independent of response modality Proc. Natl. Acad. Sci. USA 103 26 10023 10028.

    • Search Google Scholar
    • Export Citation
  • H. R. Heekeren S. Marrett L. G. Ungerleider 2008 The neural systems that mediate human perceptual decision making Nat. Rev. Neurosci. 9 6 467 479.

    • Search Google Scholar
    • Export Citation
  • S. Hochstein M. Ahissar 2002 View from the top: Hierarchies and reverse hierarchies in the visual system Neuron 36 5 791 804.

  • L. Itti C. Koch 2001 Computational modelling of visual attention Nat. Rev. Neurosci. 2 3 194 203.

  • B. Jagadeesh L. Chelazzi M. Mishkin R. Desimone 2001 Learning increases stimulus salience in anterior inferior temporal cortex of the macaque J. Neurophysiol. 86 1 290 303.

    • Search Google Scholar
    • Export Citation
  • S. Keri 2003 The cognitive neuroscience of category learning Brain Res. Brain Res. Rev. 43 1 85.

  • J. N. Kim M. N. Shadlen 1999 Neural correlates of a decision in the dorsolateral prefrontal cortex of the macaque Nat. Neurosci. 2 2 176 185.

    • Search Google Scholar
    • Export Citation
  • E. Kobatake G. Wang K. Tanaka 1998 Effects of shape-discrimination training on the selectivity of inferotemporal cells in adult monkeys J. Neurophysiol. 80 1 324 330.

    • Search Google Scholar
    • Export Citation
  • K. Koida H. Komatsu 2007 Effects of task demands on the responses of color-selective neurons in the inferior temporal cortex Nat. Neurosci. 10 1 108 116.

    • Search Google Scholar
    • Export Citation
  • Z. Kourtzi L. R. Betts P. Sarkheil A. E. Welchman 2005 Distributed neural plasticity for shape learning in the human visual cortex PLoS Biol. 3 7 e204.

    • Search Google Scholar
    • Export Citation
  • I. Kovacs P. Kozma A. Feher G. Benedek 1999 Late maturation of visual spatial integration in humans Proc. Natl. Acad. Sci. USA 96 21 12204 12209.

    • Search Google Scholar
    • Export Citation
  • C. T. Law J. I. Gold 2008 Neural correlates of perceptual learning in a sensory-motor, but not a sensory, cortical area Nat. Neurosci. 11 4 505 513.

    • Search Google Scholar
    • Export Citation
  • T. S. Lee C. F. Yang R. D. Romero D. Mumford 2002 Neural activity in early visual cortex reflects behavioral experience and higher-order perceptual saliency Nat. Neurosci. 5 6 589 597.

    • Search Google Scholar
    • Export Citation
  • R. W. Li D. M. Levi S. A. Klein 2004 Perceptual learning improves efficiency by re-tuning the decision ‘template’ for position discrimination Nat. Neurosci. 7 2 178 183.

    • Search Google Scholar
    • Export Citation
  • S. Li D. Ostwald M. Giese Z. Kourtzi 2007 Flexible coding for categorical decisions in the human brain J. Neurosci. 27 45 12321 12330.

  • W. Li V. Piech C. D. Gilbert 2004 Perceptual learning and top-down influences in primary visual cortex Nat. Neurosci. 7 6 651.

  • W. Li V. Piech C. D. Gilbert 2008 Learning to link visual contours Neuron 57 3 442 451.

  • N. K. Logothetis J. Pauls T. Poggio 1995 Shape representation in the inferior temporal cortex of monkeys Curr. Biol. 5 5 552 563.

  • R. Malach Y. Amir M. Harel A. Grinvald 1993 Relationship between intrinsic connections and functional architecture revealed by optical imaging and in vivo targeted biocytin injections in primate striate cortex Proc. Natl. Acad. Sci. USA 90 22 10469 10473.

    • Search Google Scholar
    • Export Citation
  • J. H. Maunsell S. Treue 2006 Feature-based attention in visual cortex Trends Neurosci. 29 6 317 322.

  • A. R. McIntosh M. N. Rajah N. J. Lobaugh 1999 Interactions of prefrontal cortex in relation to awareness in sensory learning Science 284 5419 1531 1533.

    • Search Google Scholar
    • Export Citation
  • B. T. Miller M. D'Esposito 2005 Searching for “the top” in top-down control Neuron 48 4 535 538.

  • E. K. Miller 2000 The prefrontal cortex and cognitive control Nat. Rev. Neurosci. 1 1 59 65.

  • E. K. Miller J. D. Cohen 2001 An integrative theory of prefrontal cortex function Annu. Rev. Neurosci. 24 167 202.

  • G. Mirabella G. Bertini I. Samengo B. E. Kilavik D. Frilli C. Della Libera L. Chelazzi 2007 Neurons in area V4 of the Macaque translate attended visual features into behaviorally relevant categories Neuron 54 2 303 318.

    • Search Google Scholar
    • Export Citation
  • Y. Miyashita 2004 Cognitive memory: cellular and network machineries and their top-down control Science 306 5695 435 440.

  • Y. Miyashita H. S. Chang 1988 Neuronal correlate of pictorial short-term memory in the primate temporal cortex Nature 331 6151 68 70.

  • C. D. Moore M. X. Cohen C. Ranganath 2006 Neural mechanisms of expert skills in visual working memory J. Neurosci. 26 43 11187 11196.

  • R. Muhammad J. D. Wallis E. K. Miller 2006 A comparison of abstract rules in the prefrontal cortex, premotor cortex, inferior temporal cortex, and striatum J. Cogn. Neurosci. 18 6 974.

    • Search Google Scholar
    • Export Citation
  • V. Navalpakkam L. Itti 2007 Search goal tunes visual features optimally Neuron 53 4 605 617.

  • K. A. Norman S. M. Polyn G. J. Detre J. V. Haxby 2006 Beyond mind-reading: Multi-voxel pattern analysis of fMRI data Trends Cogn. Sci. 10 9 424 430.

    • Search Google Scholar
    • Export Citation
  • R. M. Nosofsky 1986 Attention, similarity, and the identification-categorization relationship J. Exp. Psychol. Gen. 115 1 39 61.

  • A. J. O'Toole F. Jiang H. Abdi J. V. Haxby 2005 Partially distributed representations of objects and faces in ventral temporal cortex J. Cogn. Neurosci. 17 4 580 590.

    • Search Google Scholar
    • Export Citation
  • H. Op de Beeck J. Wagemans R. Vogels 2001 Inferotemporal neurons represent low-dimensional configurations of parameterized shapes Nature Neurosci. 4 12 1244.

    • Search Google Scholar
    • Export Citation
  • H. Op de Beeck J. Wagemans R. Vogels 2003 The effect of category learning on the representation of shape: Dimensions can be biased but not differentiated J. Exp. Psychol. Gen. 132 4 491 511.

    • Search Google Scholar
    • Export Citation
  • H. P. Op de Beeck C. I. Baker J. J. DiCarlo N. G. Kanwisher 2006 Discrimination training alters object representations in human extrastriate cortex J. Neurosci. 26 50 13025 13036.

    • Search Google Scholar
    • Export Citation
  • T. J. Palmeri I. Gauthier 2004 Visual object understanding Nature Neuroscience Reviews 5 4 291.

  • M. G. Philiastides P. Sajda 2007 EEG-informed fMRI reveals spatiotemporal characteristics of perceptual decision making J. Neurosci. 27 48 13082 13091.

    • Search Google Scholar
    • Export Citation
  • T. Poggio 1990 A theory of how the brain might work Cold Spring Harb. Symp. Quant. Biol. 55 899 910.

  • T. Poggio S. Edelman 1990 A network that learns to recognize three-dimensional objects Nature 343 6255 263 266.

  • S. Raiguel R. Vogels S. G. Mysore G. A. Orban 2006 Learning to see the difference specifically alters the most informative V4 neurons J. Neurosci. 26 24 6589 6602.

    • Search Google Scholar
    • Export Citation
  • G. Rainer H. Lee N. K. Logothetis 2004 The effect of learning on the function of monkey extrastriate visual cortex PLoS Biol. 2 2 E44.

  • G. Rainer E. K. Miller 2000 Effects of visual experience on the representation of objects in the prefrontal cortex Neuron 27 1 179 189.

    • Search Google Scholar
    • Export Citation
  • J. H. Reynolds L. Chelazzi 2004 Attentional modulation of visual processing Annu. Rev. Neurosci. 27 611 647.

  • M. Riesenhuber T. Poggio 1999 Hierarchical models of object recognition in cortex Nat. Neurosci. 2 11 1019 1025.

  • M. Riesenhuber T. Poggio 2000 Models of object recognition Nat. Neurosci. 3 Suppl 1199 1204.

  • P. R. Roelfsema 2006 Cortical algorithms for perceptual grouping Annu. Rev. Neurosci. 29 203 227.

  • P. R. Roelfsema A. van Ooyen 2005 Attention-gated reinforcement learning of internal representations for classification Neural Comput. 17 10 2176 2214.

    • Search Google Scholar
    • Export Citation
  • E. T. Rolls 1995 Learning mechanisms in the temporal lobe visual cortex Behav. Brain Res. 66 1–2 177 185.

  • E. T. Rolls N. C. Aggelopoulos F. Zheng 2003 The receptive fields of inferior temporal cortex neurons in natural scenes J. Neurosci. 23 1 339 348.

    • Search Google Scholar
    • Export Citation
  • P. Rotshtein R. N. Henson A. Treves J. Driver R. J. Dolan 2005 Morphing Marilyn into Maggie dissociates physical and identity face representations in the brain Nat. Neurosci. 8 1 107.

    • Search Google Scholar
    • Export Citation
  • M. F. Rushworth T. E. Behrens 2008 Choice, uncertainty and value in prefrontal and cingulate cortex Nat. Neurosci. 11 4 389 397.

  • K. Sakai Y. Miyashita 1991 Neural organization for the long-term memory of paired associates Nature 354 6349 152 155.

  • J. D. Schall 2001 Neural basis of deciding, choosing and acting Nat. Rev. Neurosci. 2 1 33 42.

  • C. Schiltz J. M. Bodart S. Dubois S. Dejardin C. Michel A. Roucoux M. Crommelinck G. A. Orban 1999 Neuronal mechanisms of perceptual learning: Changes in human brain activity with training in orientation discrimination Neuroimage 9 1 46 62.

    • Search Google Scholar
    • Export Citation
  • A. Schoups R. Vogels N. Qian G. Orban 2001 Practising orientation identification improves orientation coding in V1 neurons Nature 412 6846 549 553.

    • Search Google Scholar
    • Export Citation
  • D. S. Schwarzkopf Z. Kourtzi 2008 Experience shapes the utility of natural statistics for perceptual contour integration Curr. Biol. 18 15 1162 1167.

    • Search Google Scholar
    • Export Citation
  • P. G. Schyns R. L. Goldstone J. P. Thibaut 1998 The development of features in object concepts Behav. Brain Sci. 21 1 1 17.

  • M. N. Shadlen W. T. Newsome 2001 Neural basis of a perceptual decision in the parietal cortex (area LIP) of the rhesus monkey J. Neurophysiol. 86 4 1916 1936.

    • Search Google Scholar
    • Export Citation
  • D. L. Sheinberg N. K. Logothetis 2001 Noticing familiar objects in real world scenes: The role of temporal cortical neurons in natural vision J. Neurosci. 21 4 1340 1350.

    • Search Google Scholar
    • Export Citation
  • N. Sigala F. Gabbiani N. K. Logothetis 2002 Visual categorization and object representation in monkeys and humans J. Cogn. Neurosci. 14 2 187 198.

    • Search Google Scholar
    • Export Citation
  • N. Sigala N. K. Logothetis 2002 Visual categorization shapes feature selectivity in the primate temporal cortex Nature 415 6869 318 320.

    • Search Google Scholar
    • Export Citation
  • M. Sigman C. D. Gilbert 2000 Learning to find a shape Nat. Neurosci. 3 3 264 269.

  • M. Sigman G. A. Cecchi C. D. Gilbert M. O. Magnasco 2001 On a common circle: Natural scenes and Gestalt rules Proc. Natl. Acad. Sci. USA 98 4 1935 1940.

    • Search Google Scholar
    • Export Citation
  • M. Sigman H. Pan Y. Yang E. Stern D. Silbersweig C. D. Gilbert 2005 Top-down reorganization of activity in the visual pathway after learning a shape identification task Neuron 46 5 823 835.

    • Search Google Scholar
    • Export Citation
  • E. P. Simoncelli B. A. Olshausen 2001 Natural image statistics and neural representation Annu. Rev. Neurosci. 24 1193 1216.

  • M. L. Smith F. Gosselin P. G. Schyns 2004 Receptive fields for flexible face categorizations Psychol. Sci. 15 11 753 761.

  • P. L. Smith R. Ratcliff 2004 Psychology and neurobiology of simple decisions Trends Neurosci. 27 3 161 168.

  • E. Thomas M. M. Van Hulle R. Vogels 2001 Encoding of categories by noncategory-specific neurons in the inferior temporal cortex J. Cogn. Neurosci. 13 2 190.

    • Search Google Scholar
    • Export Citation
  • I. Toni M. F. Rushworth R. E. Passingham 2001 Neural correlates of visuomotor associations. Spatial rules compared with arbitrary rules Exp. Brain Res. 141 3 359 369.

    • Search Google Scholar
    • Export Citation
  • A. Treisman A. Vieira A. Hayes 1992 Automaticity and preattentive processing Am. J. Psychol. 105 2 341 362.

  • S. Treue 2003 Visual attention: The where, what, how and why of saliency Curr. Opin. Neurobiol. 13 4 428 432.

  • L. M. Vaina J. W. Belliveau E. B. des Roziers T. A. Zeffiro 1998 Neural systems underlying learning and representation of global motion Proc. Natl. Acad. Sci. USA 95 21 12657 12662.

    • Search Google Scholar
    • Export Citation
  • M. van Turennout T. Ellmore A. Martin 2000 Long-lasting cortical plasticity in the object naming system Nat. Neurosci. 3 12 1329 1334.

  • G. Wallis E. T. Rolls 1997 Invariant face and object recognition in the visual system Prog. Neurobiol. 51 2 167 194.

  • M. A. Williams S. Dang N. G. Kanwisher 2007 Only some spatial patterns of fMRI response are read out in task performance Nat. Neurosci. 10 6 685 686.

    • Search Google Scholar
    • Export Citation
  • J. M. Wolfe K. R. Cave S. L. Franzel 1989 Guided search: An alternative to the feature integration model for visual search J. Exp. Psychol. Hum. Percept. Perform. 15 3 419 433.

    • Search Google Scholar
    • Export Citation
  • T. Yang J. H. Maunsell 2004 The effect of perceptual learning on neuronal responses in monkey visual area V4 J. Neurosci. 24 7 1617 1626.

    • Search Google Scholar
    • Export Citation
  • K. Zipser V. A. Lamme P. H. Schiller 1996 Contextual modulation in primary visual cortex J. Neurosci. 16 22 7376.

Learning & Perception
Language English
Size  
Year of
Foundation
2009
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Programme
ceased
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per Year
 
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per Year
 
Publisher Akadémiai Kiadó
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H-1117 Budapest, Hungary 1516 Budapest, PO Box 245.
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Chief Executive Officer, Akadémiai Kiadó
ISSN 1789-3186 (Print)
ISSN 2060-9175 (Online)

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