Bryophyte vegetation on volcanic rock outcrops and dead wood is studied in a near-natural montane beech stand in northern Hungary. Substrate specificity of the species and the existing interspecific relationships are described. The most important species combinations and their diversity are evaluated using information theoretical functions and Monte-Carlo simulations. All analyses are based on presence/absence data of 33 species in 1508 100 cm2 microplots. Most species exhibit strong substrate specificity. Of the species that occurred with frequencies higher than 10, 8 are associated to rock, 5 to dead wood and 5 to both substrate types. Analyses of interspecific associations and agglomerative classification reveal that frequent species of species-poor bare rocks are separated from species-rich assemblages of humus-rich outcrops and coarse woody debris. Monte Carlo simulations reveal that many species combinations are significantly more frequent than expected under the assumption of random combining of species. Observed number, diversity and evenness of species combinations are significantly lower, whereas interspecific constraint (expressed as associatum) is significantly higher than under the neutral models even when data are stratified according to substrate type. The presence of coarse woody debris, not only provides habitat for wood inhabiting bryophytes, but also results in diverse rupicolous bryophyte assemblages on humus-rich outcrops.
McCullough, H.A. 1948. Plant succession on fallen logs in a virgin spruce-fir forest. Ecology 29:508-513.
'Plant succession on fallen logs in a virgin spruce-fir forest ' () 29 Ecology : 508 -513 .
McAlister, S. 1997. Cryptogam communities on fallen logs in the Duke Forest, North Carolina. J. Veg. Sci. 8:115-124.
'Cryptogam communities on fallen logs in the Duke Forest, North Carolina ' () 8 J. Veg. Sci : 115 -124 .
Kershaw, K.A. 1964. Quantitative and Dynamic Ecology. Edward Arnolds, London.
Quantitative and Dynamic Ecology , ().
Pickett, S.T.A. and P.S. White (eds). 1985. The Ecology of Natural Disturbance and Patch Dynamics. Academic Press, Orlando, Florida.
The Ecology of Natural Disturbance and Patch Dynamics , ().
Barkman, J.J. 1973. Synusial approaches to classification. In: Whittaker, R. H. (ed), Ordination and Classification of Communities. Dr. W. Junk. Publishers, The Hague, pp. 435-492.
Snusial approaches to classification , () 435 -492 .
Bartha, S. 1990. Spatial processes in developing plant communities: pattern formation detected using information theory. In: F. Krahulec, A.D.Q. Agnew, S. Agnew and J.H. Willems (eds), Spatial Processes in Plant Communities. Academia, Prague and SPB Acad. Publ., The Hague, pp. 31-47.
Satial processes in developing plant communities: pattern formation detected using information theory , () 31 -47 .
Bartha, S. 1992. Preliminary scaling for multi-species coalitions in primary succession. Abst. Bot. 16:31-41.
'Preliminary scaling for multi-species coalitions in primary succession ' () 16 Abst. Bot : 31 -41 .
Esseen, P.A., B. Ehnstróm, L. Ericson and K. Sjóberg. 1997. Boreal forests. Ecol. Bull. 46:16-47.
'Boreal forests ' () 46 Ecol. Bull : 16 -47 .
Fehér, G. and S. Orbán. 1981. A bükki óserdó korhadó fáinak mohacónológiai vizsgálata. Fol. Hist.-nat. Mus. Matr. 7:15-28.
'A bükki óserdó korhadó fáinak mohacónológiai vizsgálata ' () 7 Fol. Hist.-nat. Mus. Matr : 15 -28 .
Felfóldy, L. 1941. A debreceni Nagyerdó epiphyton vegetációja. Acta Geobot. Hung. 4:35-73.
'A debreceni Nagyerdó epiphyton vegetációja ' () 4 Acta Geobot. Hung : 35 -73 .
Gleason, H.A. 1926. The individualistic concept of the plant association. Bull. Torrey Bot. Club 53:7-26.
'The individualistic concept of the plant association ' () 53 Bull. Torrey Bot. Club : 7 -26 .
Greig-Smith, P. 1964. Quantitative Plant Ecology. Butterworth and Co. Ltd., London.
Quantitative Plant Ecology , ().
Grolle, R. 1983. Hepatics of Europe including the Azores: an annotated list of species, with synonims from the recent literature. J. Bryol. 12:403-459.
'Hepatics of Europe including the Azores: an annotated list of species, with synonims from the recent literature ' () 12 J. Bryol : 403 -459 .
Hairston, N.G., F.E. Smith, and L.B. Slobodkin. 1960. Community structure, population control, and competition. Am. Nat. 94:421-425.
'Community structure, population control, and competition ' () 94 Am. Nat. : 421 -425 .
Hübschmann, A. 1986. Prodromus der Moosgesellschaften Zentraleuropas. J. Cramer, Berlin, Stuttgart.
Prodromus der Moosgesellschaften Zentraleuropas , ().
Jonsson, B.G. 1993. The bryophyte diaspore bank and its role after small-scale disturbance in aboreal forest. J. Veg. Sci. 4:819-826.
'The bryophyte diaspore bank and its role after small-scale disturbance in aboreal forest ' () 4 J. Veg. Sci : 819 -826 .
Juhász-Nagy, P. 1976. Spatial dependence of plant populations. Part 1. Equvivalence analysis (an outline for a new model). Acta. Bot. Hung. 22:61-78.
'Spatial dependence of plant populations. Part 1. Equvivalence analysis (an outline for a new model) ' () 22 Acta. Bot. Hung : 61 -78 .
Juhász-Nagy, P. 1980. A cónológia koegzisztenciális szerkezeteinek modellezése. Thesis, MTA Budapest.
Juhász-Nagy, P. 1984. Spatial dependence of plant populations. Part 2. A family of new models. Acta. Bot. Hung. 30:363-402.
'Spatial dependence of plant populations. Part 2. A family of new models ' () 30 Acta. Bot. Hung : 363 -402 .
Harmon, M.E., J.F. Franklin, F.J. Swanson, P. Sollins, S.V. Gregory, J.D. Lattin, N.H. Anderson, S.P. Cline, N.G. Aumen, J.R. Sedell, G.W. Lienkaemper, K. Cromack, JR., and K.W. Cummins. 1986. Ecology of coarse woody debris in temperate ecosystems. Adv. Ecol. Res. 15:133-276.
'Ecology of coarse woody debris in temperate ecosystems ' () 15 Adv. Ecol. Res : 133 -276 .
Kirby, K.J., C.M. Reid, R.C. Thomas and F.B. Goldsmith. 1998. Preliminary estimates of fallen dead wood and standing dead trees in managed and unmanaged forests in Britain. J. Appl. Ecol. 35:148-155.
'Preliminary estimates of fallen dead wood and standing dead trees in managed and unmanaged forests in Britain ' () 35 J. Appl. Ecol. : 148 -155 .
Kovács, M. 1968. Die Acerion pseudoplatani Wälder (Mercurialli -Tilietum und Phyllitidi - Aceretum) des Mátra Gebirges. Acta Bot. Hung. 14:331-350.
'Die Acerion pseudoplatani Wälder (Mercurialli -Tilietum und Phyllitidi - Aceretum) des Mátra Gebirges ' () 14 Acta Bot. Hung : 331 -350 .
Lee, T.D. and G.H. La Roi. 1979. Bryophyte and understory vascular plant beta diversity in relation to moisture and elevation gradients. Vegetatio 40:29-38.
'Bryophyte and understory vascular plant beta diversity in relation to moisture and elevation gradients ' () 40 Vegetatio : 29 -38 .
Kunkel, G. P. 1975. Microhabitat and phytosociological studies of cryptogamic chasmo- and lythophytes. M. A. Thesis. Univ. of Colorado.
Microhabitat and phytosociological studies of cryptogamic chasmo- and lythophytes , ().
Ódor, P. and T. Standovár. 2001. Richness of bryophyte vegetation in a near-natural and managed beech stands: The effects of management-induced differences in dead wood. Ecol. Bull. 49:219-229.
'Richness of bryophyte vegetation in a near-natural and managed beech stands: The effects of management-induced differences in dead wood ' () 49 Ecol. Bull : 219 -229 .
Orbán, S. and L. Vajda. 1983. Magyarország mohaflórájának kézikónyve. Akadémiai Kiadó, Budapest.
Magyarország mohaflórájának kézikónyve , ().
Palmer, M.W. 1986. Pattern in corticulous bryophyte communities of the North Carolina Piedmont: Do mosses see the forest or the trees? The Bryologist 89:59-65.
'Pattern in corticulous bryophyte communities of the North Carolina Piedmont: Do mosses see the forest or the trees ' () 89 The Bryologist : 59 -65 .
Palmer, M.W. and E. van der Maarel. 1995. Variance in species richness, species association, and niche limitation. Oikos 73:203-213.
'Variance in species richness, species association, and niche limitation ' () 73 Oikos : 203 -213 .
Pentecost, A. 1980. The lichens and bryophytes of rhyolite and pumice-tuff rock outcrops in Snodownia, and some factors affecting their distribution. J. Ecol. 68:251-267.
'The lichens and bryophytes of rhyolite and pumice-tuff rock outcrops in Snodownia, and some factors affecting their distribution ' () 68 J. Ecol. : 251 -267 .
Podani, J., T. Czárán and S. Bartha. 1993. Pattern, area and diversity: the importance of spatial scale in species assemblages. Abst. Bot. 17:37-51.
'Pattern, area and diversity: the importance of spatial scale in species assemblages ' () 17 Abst. Bot : 37 -51 .
Polunin, O. and M. Walters. 1985. A Guide to the Vegetation of Britain and Europe. Oxford Univ. Press, New York.
A Guide to the Vegetation of Britain and Europe , ().
Proctor, M.C.F. 1982. Physiological ecology: water relations, light and temperature responses, carbon balance. In: A.J.E. Smith (ed.), Bryophyte Ecology. Chapman and Hall, London, pp. 333-382.
Physiological ecology: water relations, light and temperature responses, carbon balance , () 333 -382 .
Rasmussen, L. and J. Hertig. 1977. Statistical investigation of interspecific phytosociological relations in epiphytic bryophyte communities. Rev. Bryol. Lichenol. 43(2): 207-217.
'Statistical investigation of interspecific phytosociological relations in epiphytic bryophyte communities ' () 43 Rev. Bryol. Lichenol : 207 -217 .
Reice, S.R. 1994. Nonequilibrium determinants of biological community structure. Am. Sci. 82(5):424-435.
'Nonequilibrium determinants of biological community structure ' () 82 Am. Sci : 424 -435 .
Rydin, H. 1986. Competition and niche separation in Sphagnum. Can. J. Bot. 64:1817-1824.
'Competition and niche separation in ' () 64 Sphagnum. Can. J. Bot : 1817 -1824 .
Sadler, K.D. and G.E. Bradfield 2000. Microscale distribution patterns of terrestrial bryophytes in subalpine forest: the use of logistic regression as an interpretive tool. Community Ecol. 1:57-64.
'Microscale distribution patterns of terrestrial bryophytes in subalpine forest: the use of logistic regression as an interpretive tool ' () 1 Community Ecol. : 57 -64 .
Schuster, R.M. 1957. Boreal Hepaticae, a manual of the liverworts of Minnesota and adjacent regions II. Ecology. Amer. Midl. Nat. 57:203-299.
'Boreal Hepaticae, a manual of the liverworts of Minnesota and adjacent regions II. Ecology ' () 57 Amer. Midl. Nat. : 203 -299 .
Sóderstróm, L. 1988b. Niche overlap in some epixylic bryophytes. In: T. Herben and C.B. McQueen (eds.), Proceedings of the sixth CEBWG Meeting, Liblice, Czechoslovakia, pp. 108-119.
Niche overlap in some epixylic bryophytes , () 108 -119 .
Sóderstróm, L. 1988c. The occurrence of epxylic bryophyte and lichen species in an old natural and a managed forest stand in Northeast Sweden. Biol. Conserv. 45:169-178.
'The occurrence of epxylic bryophyte and lichen species in an old natural and a managed forest stand in Northeast Sweden ' () 45 Biol. Conserv : 169 -178 .
Sóderstróm, L. 1989. Regional distribution patterns of bryophyte species on spruce logs in northern Sweden. The Bryologist 92:349-355.
'Regional distribution patterns of bryophyte species on spruce logs in northern Sweden ' () 92 The Bryologist : 349 -355 .
Sóderstróm, L. 1990. Dispersal and distribution patterns in patchy, temporary habitats. In: F. Krahulec, A.D.Q. Agnew, S. Agnew and J.H. Willems (eds.), Spatial Processes in Plant Communities. SBP Publishers, The Hague, pp. 99-109.
Dispersal and distribution patterns in patchy, temporary habitats , () 99 -109 .
Standovár, T. 1998. Diversity of ground-layer vegetation in beech forest: Comparison of semi-natural and managed beech stands in Northern Hungary. In: P. Bachmann, M. Koehl and R. Paivinen (eds), Assessment of Biodiversity for Improved Forest Planning, Proceedings of the Monte Verita Conference on October 7-11, 1996. Monte Verita, Switzerland. Kluwer Academic Publishing, Dodrecht, pp. 381-388.
Diversity of ground-layer vegetation in beech forest: Comparison of semi-natural and managed beech stands in Northern Hungary , () 381 -388 .
Szollát, Gy. and S. Bartha. 1991. Pattern analysis of dolomite grassland communities using information theory models. Abst. Bot. 15:47-60.
'Pattern analysis of dolomite grassland communities using information theory models ' () 15 Abst. Bot : 47 -60 .
Vitt, D.H. and N.G. Slack. 1975. An analysis of Sphagnum-dominated kettle-hole bogs in relation to environmental gradients. Can. J. Bot. 53:332-359.
'An analysis of Sphagnum-dominated kettle-hole bogs in relation to environmental gradients ' () 53 Can. J. Bot : 332 -359 .
Vitt, D.H., H. Crum and J.A. Sneider. 1975. The vertical zonation of Sphagnum species in hummock-hollow complexes in northern Michigan. Michigan. Bot. 14:190-200.
'The vertical zonation of Sphagnum species in hummock-hollow complexes in northern Michigan ' () 14 Michigan. Bot : 190 -200 .
Watson, M. A. 1980. Patterns of habitat occupation in mosses - relevance to considerations of the niche. Bull. Torrey Bot. Club 107:346-372.
'Patterns of habitat occupation in mosses - relevance to considerations of the niche ' () 107 Bull. Torrey Bot. Club : 346 -372 .
Watt, A.S. 1947. Pattern and process in plant community. J. Ecol. 35:1-22.
'Pattern and process in plant community ' () 35 J. Ecol. : 1 -22 .
During, H.J. 1990. Clonal growth pattern among bryophytes. In: J. van Groenendael and H. de Kroon (eds.), Clonal Growth in Plants: Regulation and Function. SPB Academic Publishing, The Hague, pp. 153-176.
Clonal growth pattern among bryophytes , () 153 -176 .
During, H.J. 1992. Ecological classifications of bryophytes and lichens. In: Bates, J.W. and A.M. Farmer (eds), Bryophytes and Lichens in a Changing Environment. Clarendon Press, Oxford, pp. 1-31.
Ecological classifications of bryophytes and lichens , () 1 -31 .
Emborg, J. 1998. Undestorey light conditions and regeneration with respect to the stuctural dynamics of a near-natural temperate deciduous forest in Denmark. For. Ecol. Man. 106:83-95.
'Undestorey light conditions and regeneration with respect to the stuctural dynamics of a near-natural temperate deciduous forest in Denmark ' () 106 For. Ecol. Man : 83 -95 .
Corley, M.F.V., A.C. Crundwell, R. Düll, M.O. Hill and A.J.E. Smith. 1981. Mosses of Europe and the Azores; an annotated list of species, with synonyms from the recent literature. J. Bryol. 11:609-689.
'Mosses of Europe and the Azores; an annotated list of species, with synonyms from the recent literature ' () 11 J. Bryol : 609 -689 .
Crawley, M.J. (ed.). 1997. Plant Ecology. 2nd ed. Blackwell, Oxford.
Plant Ecology , ().
Czájlik, P. 1996. Koreloszlás és szukcesszió háborítatlan erdóallományokban: esettanulmány. In: Cs. Mátyás (ed.), Erdészeti ókológia. Mezógazdasági Kiadó, Budapest, pp. 84-92.
Koreloszlás és szukcesszió háborítatlan erdóallományokban: esettanulmány , () 84 -92 .
During, H.J. 1979. Life strategies of bryophytes: a preliminary review. Lindbergia 5:2-18.
'Life strategies of bryophytes: a preliminary review ' () 5 Lindbergia : 2 -18 .
Yarranton, G.A. and W.J. Beasleigh. 1968. Towards a mathematical model of limestone pavement vegetation. I. Vegetation and microtopography. Can. J. Bot. 46:1591-1600.
'Towards a mathematical model of limestone pavement vegetation. I. Vegetation and microtopography ' () 46 Can. J. Bot : 1591 -1600 .
Bartha, S., S.L. Collins, S.M. Glenn and M. Kertész. 1995a. Fine-scale organization of tallgrass prairie vegetation along a topographic gradient. Fol. Geobot. Phytotax. 30:169-184.
'Fine-scale organization of tallgrass prairie vegetation along a topographic gradient ' () 30 Fol. Geobot. Phytotax : 169 -184 .
Bartha, S., T. Czárán and B. Oborny 1995b. Spatial constrains masking community assembly rules: a simulation study. Folia Geobot. Phytotax. 30:471-482.
'Spatial constrains masking community assembly rules: a simulation study ' () 30 Folia Geobot. Phytotax : 471 -482 .
Bartha, S. and M. Kertész. 1998. The importance of neutral-models in detecting interspecific spatial associations from 'trainsect' data. Tiscia 31:85-98.
'The importance of neutral-models in detecting interspecific spatial associations from 'trainsect' data ' () 31 Tiscia : 85 -98 .
Bates, J. W. 1975. A quantitative investigation of the saxiculous bryophyte and lichen vegetation of Cape Clear island, county Cork. J. Ecol. 63:143-162.
'A quantitative investigation of the saxiculous bryophyte and lichen vegetation of Cape Clear island, county Cork ' () 63 J. Ecol. : 143 -162 .
Bates, J.W. and D.H. Brown. 1981. Epiphyte differentiation between Quercus petraea and Fraxinus excelsior trees in a maritime area of South West England. Vegetatio 48:61-70.
'Epiphyte differentiation between Quercus petraea and Fraxinus excelsior trees in a maritime area of South West England ' () 48 Vegetatio : 61 -70 .
Boros, Á. 1968. Bryogeographie und Bryoflora Ungarns. Akadémiai Kiadó, Budapest.
Bryogeographie und Bryoflora Ungarns , ().
Braun-Blanquet, J. 1964. Pflanzensoziologie: Grundzüge der Vegetationskunde. Springer Verlag, Wien.
Pflanzensoziologie: Grundzüge der Vegetationskunde , ().
Canham, C.D. and P.L. Marks. 1985. The response of woody plants to disturbance: patterns of establishment and growth. In: Pickett, S.T.A. and P.S. White (eds), The Ecology of Natural Disturbance and Patch Dynamics. Academic Press Inc., London, pp. 197-216.
The response of woody plants to disturbance: patterns of establishment and growth , () 197 -216 .
Clements, F.E. 1916. Plant Succession. An Analysis of the Development of Vegetation. Carnegie Institute, Washington, Publication 242, Washington D.C.
Plant Succession. An Analysis of the Development of Vegetation , ().
Collins, B.S., K.P. Dunne and S.T.A. Pickett. 1985. Responses of forest herbs to canopy gaps. In: Pickett, S.T.A. and P.S. White. (eds.), The Ecology of Natural Disturbance and Patch Dynamics. Academic Press Inc., London, pp. 218-234.
Responses of forest herbs to canopy gaps , () 218 -234 .
Corley, M.F.V. and A.C. Crundwell. 1991. Additions and amendments to the mosses of Europe and the Azores. J. Bryol. 16:337-356.
'Additions and amendments to the mosses of Europe and the Azores ' () 16 J. Bryol : 337 -356 .
Yarranton, G.A. and W.J. Baesleigh. 1969. Towards a mathematical model of limestone pavement vegetation. II. Microclimate, surface pH, and microtopography. Can. J. Bot. 47:959-974.
'Towards a mathematical model of limestone pavement vegetation. II. Microclimate, surface pH, and microtopography ' () 47 Can. J. Bot : 959 -974 .
Westhoff, V. and E. van der Maarel. 1973. The Braun-Blanquet approach. In: R.H. Whittaker (ed.), Ordination and Classification of Communities. Dr. W Junk, The Hague, pp. 617-726.
The Braun-Blanquet approach , () 617 -726 .
Yarranton, G.A. 1967. Principal component analysis of data from saxicolous bryophyte vegetation at Steps Bridge, Devon. III. Correlation of variation in the vegetation with enviromental variables. Can. J. Bot. 45:249-258.
'Principal component analysis of data from saxicolous bryophyte vegetation at Steps Bridge, Devon. III. Correlation of variation in the vegetation with enviromental variables ' () 45 Can. J. Bot : 249 -258 .
Philippi, G. 1956. Einige Moosgesellschaften des Südschwarzwal-des und den angrenzenden Rheinebene. Beitr. Nat. Forsch. Südwestdeutsch. 15:91-124.
'Einige Moosgesellschaften des Südschwarzwal-des und den angrenzenden Rheinebene ' () 15 Beitr. Nat. Forsch. Südwestdeutsch : 91 -124 .
Smith, A.J.E. 1982. Epiphytes and epiliths. In: A.J.E. Smith (ed.), Bryophyte Ecology. Champman and Hall, London, pp. 191-228.
Epiphytes and epiliths , () 191 -228 .
Sóderstróm, L. 1988a. Sequence of bryophytes and lichens in relation to substrate variables of decaying coniferous wood in Northern Sweden. Nord. J. Bot. 8:89-97.
'Sequence of bryophytes and lichens in relation to substrate variables of decaying coniferous wood in Northern Sweden ' () 8 Nord. J. Bot : 89 -97 .
Tilman, D. 1982. Resource Competition and Community Structure. Princeton Univ. Press, Princeton.
Resource Competition and Community Structure , ().
Alpert, P. 1985. Distribution quantified by microtopography in an assemblage of saxicolous mosses. Vegetatio 64:131-139.
'Distribution quantified by microtopography in an assemblage of saxicolous mosses ' () 64 Vegetatio : 131 -139 .
Barkman, J.J. 1958. Phytosociology and Ecology of Cryptogamic Epiphytes. Van Gorcum, Assen.
Phytosociology and Ecology of Cryptogamic Epiphytes , ().
Gustafsson, L. and T. Hallingback. 1988. Bryophyte flora and vegetation of managed and virgin coniferous forest in South-West Sweden. Biol. Conserv. 44:283-300.
'Bryophyte flora and vegetation of managed and virgin coniferous forest in South-West Sweden ' () 44 Biol. Conserv : 283 -300 .
Ódor, P. 2000. Description of the bryoflora and bryophyte vegetation of Kékes North Forest Reserve in Mátra Mountains (N-Hungary). Kitaibelia 5:115-123.
'Description of the bryoflora and bryophyte vegetation of Kékes North Forest Reserve in Mátra Mountains (N-Hungary) ' () 5 Kitaibelia : 115 -123 .
Podani, J. 2000. Introduction to the Exploration of Multivariate Biological Data. Backhuys Publishers, Leiden.
Introduction to the Exploration of Multivariate Biological Data , ().
McCune, B. and J.A. Antos. 1981. Correlations between forest layers in the Swan Valley, Montana. Ecology 62:1196-1204.
'Correlations between forest layers in the Swan Valley, Montana ' () 62 Ecology : 1196 -1204 .
Juhász-Nagy, P. and J. Podani. 1983. Information theory methods for the study of spatial processes and succession. Vegetatio 51:129-140.
'Information theory methods for the study of spatial processes and succession ' () 51 Vegetatio : 129 -140 .
Sjógren, E. 1964. Epilithische und epigäische Moosvegetation in Laubwäldern der Insel óland (Schweden). Ada Phyt. Suec. 48:1-184.
'Epilithische und epigäische Moosvegetation in Laubwäldern der Insel óland (Schweden) ' () 48 Ada Phyt. Suec : 1 -184 .
Slack, N.G. 1976. Host specificity of bryophytic epiphytes in eastern north America. J. Hattori Bot. Lab. 41:107-132.
'Host specificity of bryophytic epiphytes in eastern north America ' () 41 J. Hattori Bot. Lab : 107 -132 .
Podani, J. 1993. SYN-TAX-pc Computer Programs for Multivariate Data Analysis in Ecology and Systematics. Version 5.0. User's Guide. Scientia Publishing, Budapest.
SYN-TAX-pc Computer Programs for Multivariate Data Analysis in Ecology and Systematics. Version 5.0. User's Guide , ().
Slack, N.G. 1977. Species diversity and community structure in bryophytes: New York State studies. Bull. N Y. St. Mus. Sci. Surv. 428:1-70.
'Species diversity and community structure in bryophytes: New York State studies ' () 428 Bull. N Y. St. Mus. Sci. Surv : 1 -70 .