Diversity relationships of carabids in forest edges and the neighbouring forest interior and the surrounding grassland were studied. Samples were taken along three replicated forest-grassland transects using pitfall traps in the Várghis-valley (Harghita County, Romania). The study revealed significant edge effect on the carabids. Both the number of species and the Shannon diversity of carabids were significantly higher in the forest edge than in the grassland and in the forest interior. The forest was also significantly more species rich and more diverse by Shannon diversity than the grassland. Carabids of the forest interior, forest edge and grassland can be separated from each other by ordination based on abundances, suggesting that all three habitats have a distinct species assemblage. Indicator species analysis detected significant edge associated species. Our results show that the high diversity of carabids in forest edges is due both to the edge-associated species and the presence of species characteristic of the adjacent habitats.
Andersen, A. 1997. Densities of overwintering carabids and staphylinids (Col., Carabidae and Staphylinidae) in cereal and grass fields and their boundaries. J. Appl. Ent. 121:77-80.
'Densities of overwintering carabids and staphylinids (Col., Carabidae and Staphylinidae) in cereal and grass fields and their boundaries ' () 121 J. Appl. Ent. : 77 -80 .
Asteraki, E.J., C.B. Hanks and R.O. Clements 1995. The influence of different types of grassland field margin on carabid beetle (Coleoptera, Carabidae) communities. Agric. Ecosystems Environ. 54:195-202.
'The influence of different types of grassland field margin on carabid beetle (Coleoptera, Carabidae) communities ' () 54 Agric. Ecosystems Environ. : 195 -202 .
Baars, M.A. 1979. Patterns of movement of radioactive carabid beetles. Oecologia 44:125-140.
'Patterns of movement of radioactive carabid beetles ' () 44 Oecologia : 125 -140 .
Batáry, P. and Báldi, A. 2004. Evidence of an edge effect on avian net success. Conservation Biology 18:389-400.
'Evidence of an edge effect on avian net success ' () 18 Conservation Biology : 389 -400 .
Bauer, N., Z. Kenyeres and T. Kisbenedek 2004. A comparison of cluster analysis and diversity-ordering in community classification. Community Ecology 5:189-196.
'A comparison of cluster analysis and diversity-ordering in community classification ' () 5 Community Ecology : 189 -196 .
Bedford, S.E. and M.B. Usher 1994. Distribution of arthropod species across the margins of farm woodlands. Agric. Ecosystems Environ. 48:295-305.
'Distribution of arthropod species across the margins of farm woodlands ' () 48 Agric. Ecosystems Environ. : 295 -305 .
Bryan, K.M. and S.D. Wratten 1984. The responses of polyphagous predators to spatial heterogeneity: aggregation by carabid and staphylinid beetles to their cereal aphid prey. Ecol. Entomol. 9:251-259.
'The responses of polyphagous predators to spatial heterogeneity: aggregation by carabid and staphylinid beetles to their cereal aphid prey ' () 9 Ecol. Entomol. : 251 -259 .
Butovsky, R.O. 1994. Carabids in roadside ecosystems: perspectives of bioindication. In: K. Desender et al. (eds), Carabid Beetles: Ecology and Evolution. Kluwer Academic Publishers, Dordrecht, The Netherlands, pp. 241-246.
Carabids in roadside ecosystems: perspectives of bioindication , () 241 -246 .
Digby, P.G.N. and R.A. Kempton 1987. Multivariate Analysis of Ecological Communities. Chapman and Hall, London.
Multivariate Analysis of Ecological Communities , ().
Digweed, S.C., C.R. Currie, H.A. Cárcamo and J.R. Spence 1995. Digging out the digging-in effect of pitfall traps: influences of depletion and disturbance on catches of ground beetles (Coleoptera: Carabidae). Pedobiol. 39:561-576.
'Digging out the digging-in effect of pitfall traps: influences of depletion and disturbance on catches of ground beetles (Coleoptera: Carabidae) ' () 39 Pedobiol. : 561 -576 .
Dufrěne, M. and P. Legendre 1997. Species assemblages and indicator species: the need for a flexible asymmetrical approach. Ecol. Monogr. 67:345-366.
'Species assemblages and indicator species: the need for a flexible asymmetrical approach ' () 67 Ecol. Monogr. : 345 -366 .
Elek, Z., T. Magura and B. Tóthmérész 2001. Impacts of non-native Norway spruce plantation on abundance and species richness of ground beetles (Coleoptera: Carabidae). Web Ecology 2:32-37.
'Impacts of non-native Norway spruce plantation on abundance and species richness of ground beetles (Coleoptera: Carabidae) ' () 2 Web Ecology : 32 -37 .
Freude, H., K.W. Harde and G.A. Lohse 1976. Die Käfer Mitteleuropas. Goecke and Evers Verlag, Krefeld.
Die Käfer Mitteleuropas , ().
Grüm, L. 1971. Spatial differentiation of the Carabus L. (Carabidae, Coleoptera) mobility. Ekologia Polska 19:1-34.
'Spatial differentiation of the Carabus L. (Carabidae, Coleoptera) mobility ' () 19 Ekologia Polska : 1 -34 .
Holland, M.M., P.G. Risser, and R.J. Naiman 1991. Ecotones. The Role of Landscape Boundaries in the Management and Restoration of Changing Environments. Chapman and Hall, London.
Ecotones. The Role of Landscape Boundaries in the Management and Restoration of Changing Environments , ().
Horváth, R., T. Magura, G. Péter and K. Bayar 2000. Edge effect on weevil and spider communities at the Bükk National Park in Hungary. Acta Zoologica Academiae Scientiarum Hungaricae 46:275-290.
'Edge effect on weevil and spider communities at the Bükk National Park in Hungary ' () 46 Acta Zoologica Academiae Scientiarum Hungaricae : 275 -290 .
Horváth, R., T. Magura, G. Péter and B. Tóthmérész 2002. Edge effect on weevils and spiders. Web Ecology 3:43-47.
'Edge effect on weevils and spiders ' () 3 Web Ecology : 43 -47 .
Hůrka, K. 1996. Carabidae of the Czech and Slovak Republics. Kabourek, Zlin, Czech Republic.
Carabidae of the Czech and Slovak Republics , ().
Koivula, M., V. Hyyryläinen and E. Soininen 2004. Carabid beetles (Coleoptera: Carabidae) at forest-farmland edges in southern Finland. Journal of Insect Conservation 8:297-309.
'Carabid beetles (Coleoptera: Carabidae) at forest-farmland edges in southern Finland ' () 8 Journal of Insect Conservation : 297 -309 .
Lövei, G. L. 2005. Generalised entropy indices have a long history in ecology - a comment. Community Ecology 6:245-247.
'Generalised entropy indices have a long history in ecology - a comment ' () 6 Community Ecology : 245 -247 .
Lövei, G. and K.D. Sunderland 1996. Ecology and behavior of ground beetles (Coleoptera: Carabidae). Ann. Rev. Entomol. 41:231-256.
'Ecology and behavior of ground beetles (Coleoptera: Carabidae) ' () 41 Ann. Rev. Entomol. : 231 -256 .
Lövei, G. L., T. Magura, B. Tóthmérész and V. Ködöböcz. 2006. The influence of matrix and edges on species richness patterns of ground beetles (Coleoptera, Carabidae) in habitat islands. Global Ecology and Biogeography 15: 283-289.
'The influence of matrix and edges on species richness patterns of ground beetles (Coleoptera, Carabidae) in habitat islands ' () 15 Global Ecology and Biogeography : 283 -289 .
Magura, T. 1995. Wanderungsverhalten und Ortstreue der Arten Abax ater und Abax ovalis (Coleoptera: Carabidae). Folia Entomologica Hungarica 56:89-99.
'Wanderungsverhalten und Ortstreue der Arten Abax ater und Abax ovalis (Coleoptera: Carabidae) ' () 56 Folia Entomologica Hungarica : 89 -99 .
Magura, T. and B. Tóthmérész 1997. Testing edge effect on carabid assemblages in an oak-hornbeam forest. Acta Zool. Acad. Sci. Hung. 43:303-312.
'Testing edge effect on carabid assemblages in an oak-hornbeam forest ' () 43 Acta Zool. Acad. Sci. Hung. : 303 -312 .
Magura, T. and B. Tóthmérész 1998. Edge effect on carabids in an oak-hornbeam forest at the Aggtelek National Park (Hungary). Acta Phytopathol. Entomol. Hung. 33:379-387.
'Edge effect on carabids in an oak-hornbeam forest at the Aggtelek National Park (Hungary) ' () 33 Acta Phytopathol. Entomol. Hung. : 379 -387 .
Magura, T., B. Tóthmérész and Zs. Bordán 2000a. Effects of nature management practice on carabid assemblages (Coleoptera: Carabidae) in a non-native plantation. Biol. Conserv. 93:95-102.
'.Effects of nature management practice on carabid assemblages (Coleoptera: Carabidae) in a non-native plantation ' () 93 Biol. Conserv. : 95 -102 .
Magura, T., B. Tóthmérész and Z. Elek 2003. Diversity and composition of carabids during a forestry cycle. Biodiversity and Conservation 12:73-85.
'Diversity and composition of carabids during a forestry cycle ' () 12 Biodiversity and Conservation : 73 -85 .
Magura, T., B. Tóthmérész and Z. Elek 2004. Effects of leaf-litter addition on carabid beetles in a non-native Norway spruce plantation. Acta Zoologica Academiae Scientarium Hungaricae 50:9-23.
'Effects of leaf-litter addition on carabid beetles in a non-native Norway spruce plantation ' () 50 Acta Zoologica Academiae Scientarium Hungaricae : 9 -23 .
Magura, T., B. Tóthmérész and T. Molnár 2000b. Spatial distribution of carabid species along grass-forest transects. Acta Zool. Acad. Sci. Hung. 46:1-17.
'.Spatial distribution of carabid species along grass-forest transects ' () 46 Acta Zool. Acad. Sci. Hung. : 1 -17 .
Magura, T., B. Tóthmérész and T. Molnár 2001a. Edge effect on carabids along forest-grass transects. Web Ecology 2:7-13.
'Edge effect on carabids along forest-grass transects ' () 2 Web Ecology : 7 -13 .
Magura, T., B. Tóthmérész and T. Molnár 2001b. Forest edge and diversity: carabids along forest-grassland transects. Biodiversity and Conservation 10:287-300.
'Forest edge and diversity: carabids along forest-grassland transects ' () 10 Biodiversity and Conservation : 287 -300 .
Molnár, T., T. Magura, B. Tóthmérész and Z. Elek 2001. Ground beetles (Carabidae) and edge effect in oak-hornbeam forest - grassland transects. European Journal of Soil Biology 37:297-300.
'Ground beetles (Carabidae) and edge effect in oak-hornbeam forest - grassland transects ' () 37 European Journal of Soil Biology : 297 -300 .
Murcia, C. 1995. Edge effects in fragmented forests: implications for conservation. Trends Ecol. Evol. 10:58-62.
'Edge effects in fragmented forests: implications for conservation ' () 10 Trends Ecol. Evol. : 58 -62 .
Niemelä, J. 1988. Carabid beetles in shore habitats on the Åland islands, SW Finland: the effect of habitat availability and species characteristics. Acta Oecol. 9:379-395.
'Carabid beetles in shore habitats on the Åland islands, SW Finland: the effect of habitat availability and species characteristics ' () 9 Acta Oecol. : 379 -395 .
Ricotta, C. 2005. On parametric diversity indices in ecology: A historical note. Community Ecology 6:241-244.
'On parametric diversity indices in ecology: A historical note ' () 6 Community Ecology : 241 -244 .
Samways, M. J. 2005. Insect Diversity Conservation. Cambridge Univ. Press, Cambridge.
Insect Diversity Conservation , ().
Spence, J.R. and J. Niemelä 1994. Sampling carabid assemblages with pitfall traps: the madness and the method. Can. Ent. 126:881-894.
'Sampling carabid assemblages with pitfall traps: the madness and the method ' () 126 Can. Ent. : 881 -894 .
Spence, J.R., D.W. Langor, J. Niemelä, H.A. Cárcamo and C.R. Currie 1996. Northern forestry and carabids: the case for concern about old-growth species. Ann. Zool. Fennici 33:173-184.
'Northern forestry and carabids: the case for concern about old-growth species ' () 33 Ann. Zool. Fennici : 173 -184 .
Taboada, A., D.J. Kotze and J.M. Salgado 2004. Carabid beetle occurrence at the edges of oak and beech forests in NW Spain. Eu. J. Entomol. 101:555-563.
'Carabid beetle occurrence at the edges of oak and beech forests in NW Spain ' () 101 Eu. J. Entomol. : 555 -563 .
Thiele, H.U. 1977. Carabid Beetles in their Environments. Springer Verlag, Berlin.
Carabid Beetles in their Environments , ().
Tóthmérész, B. 1995. Comparison of different methods for diversity ordering. J. Veg. Sci. 6:283-290.
'Comparison of different methods for diversity ordering ' () 6 J. Veg. Sci. : 283 -290 .
Tóthmérész, B. 1998. On the characterization of scale-dependent diversity. Abstracta Botanica 22:149-156.
'On the characterization of scale-dependent diversity ' () 22 Abstracta Botanica : 149 -156 .
Tóthmérész, B. 2005. Diversity Characterizations. In R. In: L.L Gövei and S. Toft (eds.), European Carabidology 2003. Proceedings of the 11th European Carabidologists' Meeting, DIAS Report No. 114, Flakkebjerg. pp. 333-344.
'Diversity Characterizations ' , , .
Tóthmérész, B. and T. Magura 2005a. Affinity indices for environmental assessment using carabids. In: L.L.G.övei and S. Toft (eds.), European Carabidology 2003. Proceedings of the 11th European Carabidologists' Meeting. DIAS Report No. 114, Flakkebjerg. pp. 345-352.
, , .
Tóthmérész, B. and T. Magura 2005b. Diversity and scalable diversity characterizations. In: L.LGövei and S. Toft (eds.), European Carabidology 2003. Proceedings of the 11th European Carabidologists' Meeting. DIAS Report No. 114, Flakkebjerg. pp. 353-368.
, , .
Wallin, H. 1986. Habitat choice of some field-inhabiting beetles (Coleoptera: Carabidae) studied by recapture of marked individuals. Ecol. Entomol. 11:457-466.
'Habitat choice of some field-inhabiting beetles (Coleoptera: Carabidae) studied by recapture of marked individuals ' () 11 Ecol. Entomol. : 457 -466 .
Zar, J.H. 1984. Biostatistical Analysis. Prentice Hall International, London.
Biostatistical Analysis , ().