Introduction

This article continues the study of the taxonomy of the genus Chrysolina Motschulsky, 1860 in the mountainous regions of China, begun in previous works (Bieńkowski 2022, 2023, 2024). The present article is devoted to the designation of a new subgenus оf the genus Chrysolina which includes six previously known species and 14 new ones from Hengduan Mountains (Sichuan and Yunnan Provinces).

All previously described species, which I assign here to a new subgenus, were described without inclusion in any subgenus of the genus Chrysolina. The first two species of the subgenus in question, namely Ch. nixiana Lopatin, 2008 and Ch. yunnana Lopatin, 2008, were described by Lopatin (2008). Subsequently Daccordi et al. (2011) described Ch. boccaccioi Daccordi et Yang, 2011, Ch. luobinwangi Ge et Daccordi, 2011, and Ch. wangboi Daccordi et Yang, 2011. Ge et al. (2011) described Ch. hongyuanensis Daccordi et Ge, 2011.

Bieńkowski (2019) established unnamed species group that included Ch. boccaccioi and Ch. wangboi, as well as another unnamed species group that included Ch. nixiana and Ch. yunnana, and compiled keys to the species of these two groups; species Ch. luobinwangi was considered to be close to two unrelated species Ch. claripes Lopatin, 2002 and Ch. lijieae Daccordi et Yang, 2011, and Ch. hongyuanensis was considered separately from the others.

In the present article, based on the study of comparative morphology, it is established that the six species under consideration as well as 14 new species form a natural group that has been assigned the status of the new subgenus Latipoda.

Materials and methods

Materials, including the type ones, from the following museums (names of curators in brackets) and private collections have been studied:

All holotypes and some paratypes of the new species are deposited in ZIN. The other paratypes are deposited in BC, DC, RC, IZCAS, and USNM. During the present study, the type specimens of all known species were examined.

Morphological description and terminology of structural details mostly follow Bieńkowski (2019), with the addition of special term relating to prothoracic hypomeron, namely anterior cavity (Bieńkowski 2024) (Fig. 1F a). The microrelief (visible at magnification of about 60 X) of the dorsal body surface usually includes microreticulation (microcells) and/or micropunctules in the places of connection of microcells (Fig. 2D). The diameter of hollow-shaped punctures of elytron is measured at the bottom of the puncture (Fig. 2E a). Schemes for measuring the antennomeres, palpomeres and tarsomeres are shown in Fig. 1G, H. Antennomeres and palpomeres are not cylindrical, but slightly flattened dorso-ventrally, and the measurements are taken along the wider side. The length of the aedeagus is the straight line distance between the base and the apex, the other measurements are shown in Fig. 3D a, b. Metric characters were studied under a stereomicroscope using a measuring eyepiece. The scale division value when measuring the diameter of punctures is 0.01 mm, for other measurements – 0.1 mm.

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Chrysolina species: A–E − Ch. nixiana, male aedeagus, dorsal and lateral view: A – holotype, B–E − variabilty of the specimens from one locality (Yunnan, Tianbaoshan); F–H – Ch. nixiana, male: F – left prothoracic hypomeron, ventral view (a – place of anterior cavity, b – place of basal fold), G – maxillary palpus (measurements: a – length of 4^th palpomere, b – width of 4^th palpomere, c – length of 3^rd palpomere, d – width of 3^rd palpomere), H – 1^st fore-tarsomere, dorsal view (measurements: a – width, b – length); I – Ch. circe sp. nov., holotype, male aedeagus, dorsal and lateral view; J–K – male aedeagus, dorsal, ventral, and lateral view: J – Ch. margaritifera sp. nov., holotype; K– Ch. luluni sp. nov., holotype

Citation: Animal Taxonomy and Ecology 2024; 10.1556/1777.2024.00035

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Chrysolina species: A–B – Ch. patrikeevi sp. nov., holotype, aedeagus, dorsal and lateral view; C – Ch. margaritifera sp. nov., holotype, male, relief of elytron; D – scheme of elytral microrelief with cells and micropunctules

Citation: Animal Taxonomy and Ecology 2024; 10.1556/1777.2024.00035

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Chrysolina species, male aedeagus: A–C – Ch. konstantinovi sp. nov.: A – holotype, B–C – paratypes (B – male from the type locality, C – male from Hutiao gorge); D – Ch. pakhomovae sp. nov., holotype (measurements: a – length of cover plate, b – length of apical orifice); E − Ch. selene sp. nov., holotype; F – Ch. leda sp. nov., holotype. A–E − dorsal and lateral view, F – dorsal, ventral, and lateral view

Citation: Animal Taxonomy and Ecology 2024; 10.1556/1777.2024.00035

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The original descriptions of previously described species were incomplete and did not contain a number of important diagnostic features. Therefore, all these species are redescribed. The species descriptions are ordered as follows: previously described species first, alphabetically, then new species, alphabetically.

Photographs of the beetles were made by a Nikon D-90 digital camera, combined with Tamron SP 70–300 mm F/4-5.6 and inverted Vega-12B 2.8-90 lenses, using Helicon Remote 3.9.10 W software. Images of the same objects at different focal planes were combined using Helicon Focus 6 software.

Taxonomy

Family Chrysomelidae

Genus Chrysolina Motschulsky, 1860

Type species: Chrysomela staphylaea Linnaeus, 1758, by original designation

Chrysolina (Latipoda subgen. nov.)

https://zoobank.org/0FE59661-2490-40CD-AE3D-E82E5BAAD6EC

Diagnosis. Body usually dark metallic, mostly bronze, rarely violet or green, or black, moderately shining or sericeous, but not dull.

Last maxillary palpomere narrow, barrel-shaped, never securiform, as wide as penultimate one or slightly broader than the latter, similar in both sexes (in the species for which both sexes are known).

Pronotum without anterior and posterior setiferous pores. Pronotal lateral impression broad, shallow to obsolete, never furrow-shaped, usually with more or less large punctures. Anterior side marginated, ciliate. Posterior side entirely marginated.

Prothoracic hypomeron without basal fold, with anterior cavity small or absent. Anterior coxal cavities broadly opened behind. Metasternum marginated anteriorly.

Humeral callus of elytron obsolete or absent. Elytral punctures arranged in rows completely regular or partly irregular. Elytral intervals flat or slightly convex. Sutural furrow developed at apical slope. Scutellar puncture row present, but sometimes reduced to 1–2 punctures.

Elytral epipleura inclined outside, visible along entire length in lateral view, apically with setae more or less numerous.

Hind wings absent.

Pygidium without impression or furrow in apical ½, with weak impression in basal ½.

Last abdominal ventrite simple, convex in both sexes.

1^st–3^rd tarsomeres more or less broadened in male, narrow in female, with entire sole in both sexes (in the species for which both sexes are known). Claw tarsomere without denticles beneath. Claws simple and separate from each other.

Aedeagus tube-shaped, more or less depressed (in lateral view), with simple apical margin, with apical denticles in some species, with flagellum tube-shaped, simple, exposed, rarely absent.

Differential diagnosis. The new subgenus belongs to the wingless mountain Chrysolina groups in the subtropics of China. It differs from the subgenera whose representatives inhabit the mountains of Sichuan Province and the adjacent part of Yunnan Province in the following ways.

It differs from the subgenus Medvedevlevna Özdikmen, 2008 in the last maxillary palpomere longer than broad, longer than penultimate one or as long as the latter, elytral punctures arranged in rows, while Medvedevlevna has last maxillary palpomere broader than long or as long as broad, rather shorter than penultimate one, and elytral punctures entirely irregular.

It differs from Ch. altimontana (Rybakow, 1889) species group of the subgenus Pezocrosita Jacobson, 1901 in the absence of long glabrous basal “heel” at 1^st tarsomere, while Ch. altimontana species group has 1^st tarsomere bearing long narrow glabrous basal “heel” ventrally.

It differs from Ch. obovata (Jacobson, 1895) species group of the subgenus Pezocrosita Jacobson, 1901 in the absence of narrow pronotal lateral furrow in basal ¼–½.

It differs from the subgenus Semenowia Weise, 1889 in last maxillary palpomere narrow, barrel-shaped, similar in both sexes, and the absence of narrow deep pronotal lateral furrow in basal ½, while Semenowia has male last maxillary palpomere broad, securiform, broader than those in female.

It differs from the subgenus Timarchomela Achard, 1922 in dorsum more shining, pronotum with lateral impression shallow, but distinct, elytral punctures always distinct, while Timarchomela has dorsum dull or sericeous, pronotum without distinct lateral impression, elytral punctures mostly obsolete, hardly visible (except Ch. costulata (Achard, 1922)).

It differs from Ch. (Allohypericia Bechyné, 1950) mauroi Lopatin, 2005 in the pronotal lateral impression not deepened in basal ⅓.

It differs from the subgenus Volosatik Bieńkowski, 2022 in the body less shining, not brightly colored, but dark metallic (mostly bronze) or black, moderately shining or sericeous; pronotal lateral impression usually with more or less large punctures, male 1^st–3^rd tarsomeres more or less broadened, broader than those in female. All members of Volosatik inhabit NW part of Yunnan far from the area of the new subgenus.

Type species: Chrysolina nixiana Lopatin, 2008

Distribution: China: eastern slope of Hengduan Mountains (Sino-Tibetan mountains) in Sichuan Province and adjacent areas of Yunnan Province at altitudes mostly in the alpine meadow region from 3,000 to 4,915 m above sea level, only one species (Ch. konstantinovi sp. nov.) – from 2,000 to 3,500 m.

Material examined: Holotype: China, W Sichuan Province, road Sabdè–Jiulong, pass 40 km N of Jiulong, alpine region, 3,500 m, 22–23.6.1994, J. Kaláb leg.: male (DC); paratypes: China, W Sichuan Province, road Sabdè–Jiulong, pass 40 km N of Jiulong, alpine region: 1 female (BC); China, W Sichuan Province, road Sabdè–Jiulong, pass 40 km N of Jiulong, alpine region, 3,500 m, 22.6.1994, J. Kaláb leg.: 1 female (NME), 1 female (NMB).

Description. Body elongate oval, 1.7 X as long as wide (Fig. 5A). Body black with trochanters and underside of 1^st and 2^nd antennomeres rufous. Dorsum moderately shining, microreticulate, besides that head and pronotum micropunctulate.

Supraorbital line deep, long, but not reaching antennal insertion. Base of antenna 2.8 X closer to clypeus than to eye. Last maxillary palpomere narrow, barrel-shaped, obliquely truncate apically, 1.3 X as long as wide, 1.1 X as long as penultimate palpomere, as wide as the latter, similar in both sexes. Antenna 0.4 X as long as body. 7^th antennomere 0.9 X as long as wide, antennomere 9 1.1 X as long as wide.

Pronotum 1.9–2.0 X as wide as long, broadest at mid-length, with about the same width toward base, with lateral sides almost straight and parallel in basal ½ and rounded in anterior ½, with anterior side moderately emarginate, anterior angles roundly triangular. Lateral callus very weakly convex, almost flat along entire length; lateral impression very broad, obsolete. Pronotal disc unevenly covered with mixed large (0.06 mm), moderate (0.03 mm), and fine (0.02 mm) punctures. Lateral impression with numerous large (0.06 mm) punctures and wrinkled intervals. Large punctures along outer margin of lateral callus absent. Pronotum entirely marginate and ciliate anteriorly, entirely marginate posteriorly. Anterior and posterior setiferous pores of pronotum absent.

Prothoracic hypomeron weakly convex, with weak impression and weak transverse wrinkles laterally. Anterior cavity and basal fold of prothoracic hypomeron absent.

Metasternum entirely marginate anteriorly.

Elytron without humeral callus. Elytral punctures moderate to large (0.03–0.05 mm), hollow-shaped, arranged in regular, slightly paired rows, with interspaces between punctures slightly convex, rugose, covered by fine (0.02 mm) punctures. 5^th row with 15 punctures. Scutellar row with 4 punctures. Sutural furrow very long, distinct along entire length of apical slope. Elytral epipleuron inclined outside, visible along entire length in lateral view, with only few setae apically.

Hind wing absent.

1^st–3^rd tarsomeres with entire sole in both sexes, narrow.

Pygidium with shallow longitudinal impression in basal ½ only in both sexes.

Last abdominal ventrite simple, convex in both sexes.

Aedeagus (Fig. 4A) tube-shaped, with parallel lateral sides at most of its length, rectilinearly narrowed at sides of apical orifice, and bearing trapeziform apical projection. Aedeagus without apical denticles. Aedeagus strongly curved in lateral view, with apical part recurved upward. Flagellum tube-shaped, simple, exposed. Apical orifice 0.2 X as long as aedeagus length, cover plate 0.2 X as long as apical orifice.

Female spermatheca present, but rudimentary, whip-shaped.

Material examined: Paratypes: China, N Sichuan Province, Hongyuan, 4,200 m, 21.7–3.8.1991, J. Kaláb leg.: 1 male (BC), 1 male (NME), 1 male (DC), 1 male (NMB). Non-type specimen: China, Sichuan Province, Ngawa Tibetan and Quiang Autonomous Pref., Shuamalukou pass, 31°57′ N/102°38′ E, V. Patrikeev leg.: 1 male (RC).

Description. Male (Fig. 5B). Body oval, 1.6 X as long as wide. Dorsal side dark bronze; underside and legs black with bronze reflection; antennae piceous with 1^st and 2^nd antennomeres rufous below apically, base of trochanters reddish. Dorsum moderately shining, microreticulate and micropunctulate.

Supraorbital line distinct, long, but not reaching antennal base. Base of antenna 2.3 X closer to clypeus than to eye. Last maxillary palpomere narrow, barrel-shaped, obliquely truncate at apex, 1.6 X as long as wide, 1.3 X as long as penultimate palpomere and 1.1 X as wide as the latter. Antenna 0.5 X as long as body. 7^th antennomere 1.3 X as long as wide, 9^th antennomere 1.5 X as long as wide.

Pronotum 2 X as wide as long, broadest before base, with lateral sides rounded, more convergent forward than backward, with anterior side moderately emarginate, anterior angles slightly rounded. Lateral callus moderately convex along entire length; lateral impression broad and very shallow along entire length. Pronotal disc evenly covered with dense, fine (0.02 mm) punctures. Lateral impression with numerous (about 35–40) moderate (0.03 mm) punctures. Several moderate (0.03 mm) punctures present also along outer margin of lateral callus. Pronotum entirely marginate and ciliate anteriorly, entirely marginate posteriorly. Anterior and posterior setiferous pores of pronotum absent.

Prothoracic hypomeron almost flat, without lateral impression, with obsolete irregular wrinkles laterally. Anterior cavity of prothoracic hypomeron small, basal fold absent.

Metasternum entirely marginate anteriorly.

Elytron without humeral callus. Elytral punctures moderate (0.03 mm), partly arranged in regular, some undulate, paired (2–3, 4–5, 6–7, 8–9) rows, with intervals flat. Broad intervals between pairs of rows covered with numerous fine (0.02 mm) punctures. Intervals with numerous obsolete wrinkles. 5^th row with about 40 punctures. Scutellar row with 8–9 punctures. Sutural furrow distinct at apical slope. Elytral epipleuron inclined outside, visible along entire length in lateral view, with dense setae apically.

Hind wing absent.

1^st–3^rd tarsomeres, especially 1^st tarsomere, moderately broadened, with entire sole. 1^st fore-tarsomere as wide as long, as wide as 3^rd, 2.9 X as wide as claw tarsomere.

Pygidium with shallow, obsolete longitudinal impression in basal ½ only.

Last abdominal ventrite simple, convex, with apical margin broadly emarginate and bearing furrow along apical margin.

Aedeagus (Figs 7F and 8A) flattened dorso-ventrally, 3.9 X as long as wide, slightly broadened in apical ½, then rectilinearly narrowed at sides of apical orifice, with apical margin anchor-shaped. Aedeagus with large apical denticles ventrally. Aedeagus moderately curved in lateral view. Flagellum simple, tube-shaped, slightly exposed. Apical orifice 0.2 X as long as aedeagus length, cover plate 0.3 X as long as apical orifice. Pores present on dorsal, lateral and ventral sides of apical margin.

Body length: 6.6–7.1 mm; aedeagus 2.7–2.8 mm long, 0.7 mm wide.

Description of female. 3^rd hind tarsomere with entire sole (Ge et al. 2011).

Chrysolina luobinwangi Ge et Daccordi in: Daccordi et al. 2011: 304

Material examined: paratype: China, Sichuan Province, environs of Kangding, Mogecuo Nat. Park, 3,000–3,200 m, 12–15.8.1996: 1 female (BC). Non type specimen (topotype): China, Sichuan Province, Tagu, 07.1992: 1 male (BC).

Description. Body broadly oval, 1.6 X as long as wide (Fig. 5C). Dorsal side dark bronze; underside and legs black with weak bluish or greenish metallic reflection; base of trochanters rufous; antennae piceous with 1^st and 2^nd antennomeres rufous below. Dorsum moderately shining (male) or sericeous (female), microreticulate and micropunctulate.

Supraorbital line distinct above eye. Base of antenna 1.8 X closer to clypeus than to eye. Last maxillary palpomere narrow, barrel-shaped, obliquely truncate apically, 1.3 X as long as wide, 1.5 X as long as penultimate palpomere, 1.3 X as wide as the latter, similar in both sexes. Antenna 0.4 X as long as body. 7–9^th antennomeres as long as broad.

Pronotum 1.9 X as wide as long, broadest basally, with lateral sides almost straight and slightly convergent in basal ½ and rounded in anterior ½, with anterior side moderately emarginate, anterior angles rounded. Lateral callus weakly convex along entire length; lateral impression broad, very weak, slightly more developed at base and in anterior ½. Pronotal disc evenly covered with dense, moderate (0.03 mm) punctures. Lateral impression with numerous (18–25) large (0.05–0.06 mm) punctures. Large punctures along outer margin of lateral callus absent. Pronotum entirely marginate and ciliate anteriorly, entirely marginate posteriorly. Anterior and posterior setiferous pores of pronotum absent.

Prothoracic hypomeron weakly impressed and covered with irregular wrinkles laterally. Anterior cavity and basal fold of prothoracic hypomeron absent.

Metasternum entirely marginate anteriorly.

Elytron without humeral callus. Elytral punctures moderate (0.03 mm), mostly arranged in regular, equidistant rows, with intervals slightly convex, covered by fine wrinkles. 5^th row with 16–21 punctures. Scutellar row with 4 punctures. Sutural furrow distinct at apical slope. Elytral epipleuron inclined outside, visible along entire length in lateral view, sparsely ciliate apically.

Hind wing absent.

1^st–3^rd tarsomeres very broad, with entire sole in male. Male 1^st fore-tarsomere 1.4 X as wide as long, 0.9 X as wide as 3^rd tarsomere, 2.8 X as wide as claw tarsomere. Female 3^rd hind-tarsomere with entire sole (Daccordi et al. 2011).

Pygidium with weak, shallow longitudinal impression in basal ½ only in both sexes.

Last abdominal ventrite simple, convex in both sexes, with apical margin broadly truncate in male, rounded in female.

Aedeagus (Figs 4E and 8B) 4.3 X as long as wide, with almost parallel lateral sides, hardly broadened toward apex, with apical margin roundly triangular. Aedeagus without apical denticles. Aedeagus moderately evenly curved in lateral view. Flagellum narrow, simple, exposed. Apical orifice 0.3 X as long as aedeagus length, cover plate 0.4 X as long as apical orifice. Pores present along lateral side of apical orifice (viewed laterally).

Female spermatheca absent (Daccordi et al. 2011).

Body length: 6.2 mm (male), 7.4 mm (female); aedeagus 2.4 mm long, 0.6 mm wide.

Variability (Daccordi et al. 2011). Male 5.4 mm long, female 7.8 mm long. In the male (holotype), antennae and legs reddish brown; in the available specimens, antennae piceous, legs black with weak metallic reflection.

Chrysolina nixiana Lopatin, 2008: 837

Material examined: Holotype: China, N Yunnan Province, S of Nixi Village, 4,085 m, 27°55′14″N / 99°31′18″E, 17.07.2007, I. Belousov, I. Kabak leg.: male (ZIN). Non type specimens: China, Yunnan Province, Tianbaoshan between Shangrila & Habaxue Shan, NE of Xiaozhongdian, 4,160 m, 27°35′47″ N / 99°54′34″ E, 21.5.2015, I. Belousov, I. Kabak, G. Davidian leg.: 2 males, 2 females (BC); China, Yunnan Province, Tianbaoshan between Shangrila & Habaxue Shan, NE of Xiaozhongdian, 4,015 m, 27°36′0″ N / 99°54′5″ E, 20.5.2015, I. Belousov, I. Kabak, G. Davidian leg.: 15 males, 12 females (BC), 4 males (RC).

Description. Body elongate oval, 1.7 X as long as wide (Fig. 5D). Body one-colored dark bronze or head and pronotum greenish bronze, elytra coppery, with underside of 1^st and mostly 2^nd antennomeres rufous. Dorsum entirely moderately shining (male) or with sericeous elytra (female), microreticulate and micropunctulate.

Supraorbital line deep, long, but not reaching antennal insertion. Base of antenna 1.7–1.8 X closer to clypeus than to eye. Last maxillary palpomere narrow, barrel-shaped, transversely truncate apically, 1.5–1.7 X as long as wide, 1.3–1.5 X as long as penultimate palpomere, 1.1–1.2 X as wide as the latter, similar in both sexes (Fig. 1G). Antenna 0.4 X as long as body. 7^th antennomere 1.6–1.9 X as long as wide, 9^th antennomere 1.7–1.9 X as long as wide.

Pronotum 1.8–1.9 X as wide as long, broadest at mid-length, with lateral sides rounded, more convergent forward than backward, with anterior side deeply emarginate, anterior angles rounded. Lateral callus weakly convex along entire length; lateral impression very broad, shallow. Pronotal disc evenly covered with moderate (0.03 mm) punctures or covered with fine (0.02 mm) punctures with some moderate punctures mostly laterally. Lateral impression with numerous (about 25) large (0.06–0.09 mm) punctures. Moderate (0.03 mm) punctures present along outer margin of lateral callus. Pronotum entirely marginate and ciliate anteriorly, entirely marginate posteriorly. Anterior and posterior setiferous pores of pronotum absent.

Prothoracic hypomeron (Fig. 1F) weakly convex, with shallow impression covered by distinct or obsolete wrinkles laterally. Anterior cavity of hypomeron small or absent, basal fold absent.

Metasternum entirely marginate anteriorly.

Elytron without humeral callus. Elytral punctures moderate to large (0.03–0.06 mm), hollow-shaped, arranged in regular, slightly paired rows, more or less visible among punctures in intervals. Scutellar puncture row present. Interspaces between punctures slightly convex, rugose, covered by fine (0.02 mm) punctures. 5^th row with 13–18 punctures. Sutural furrow very long, distinct at entire length of apical slope. Elytral epipleuron inclined outside, visible along entire length in lateral view, with sparse setae apically.

Hind wing absent.

1^st–3^rd tarsomeres with entire sole in both sexes, narrow in female, very broad in male. Male 1^st fore-tarsomere 1.0–1.1 X as wide as long, as wide as 3^rd tarsomere, 4.4–4.5 X as wide as claw tarsomere (Fig. 1H).

Pygidium with shallow longitudinal impression in basal ½ only in both sexes.

Last abdominal ventrite simple, convex and slightly swollen, with apical margin broadly truncate and bearing marginal furrow in both sexes.

Aedeagus (Figs 1A–E, 8C) flattened dorso-ventrally, with dorsal side concave along most of its length, 5.2 X as long as wide, with parallel lateral sides at most of its length, more or less narrowed at sides of apical orifice, and bearing roundly trapeziform apical projection. Aedeagus without apical denticles. Aedeagus strongly curved, S-shaped in lateral view. Flagellum narrow, simple, exposed. Apical orifice 0.6 X as long as aedeagus length, cover plate 0.2 X as long as apical orifice. Pores present on lateral and ventral sides apically. Shape of apical part variable (Fig. 1A–E).

Female spermatheca present.

Body length: 7.1–7.6 mm (male), 7.6–8.5 mm (female); aedeagus 3.1 mm long, 0.6 mm wide.

Chrysolina wangboi Daccordi et Yang in: Daccordi et al. 2011: 311

Material examined: Paratypes: China, Sichuan Province, Zheduo Shan, 35 km N of Kanding, 10.7.2002, Beneš leg.: 1 male (DC), 1 male (BC); China, Sichuan Province, environs of Kangding, Modecuo Nat. Park, 3,000–3,200 m, 12–15.8.1996: 1 female (BC); China, W Sichuan Province, Zheduo Shan, W of Kanding, alpine region with Rhododendron, 4,300–4,500 m, 12–22.6.2004, R. Fabbri leg.: 1 female (BC).

Description. Body elongate oval, 1.7 X as long as wide (Fig. 10A). Body black with underside of 1^st and 2^nd antennomeres rufous. Dorsum moderately shining, microreticulate and micropunctulate.

Supraorbital line deep, long, but not reaching antennal insertion. Base of antenna 2.3 X closer to clypeus than to eye. Last maxillary palpomere narrow, barrel-shaped, obliquely truncate apically, 1.4 X as long as wide, 1.1 X as long as penultimate palpomere, as wide as the latter, similar in both sexes. Antenna 0.4 X as long as body. 7^th antennomere 1.5 X as long as wide, 9^th antennomere 1.3 X as long as wide.

Pronotum 2.1 X as wide as long, broadest at base, with lateral sides convergent forward, almost straight in basal ½ and rounded in anterior ½, with anterior side deeply emarginate, anterior angles roundly triangular. Lateral callus very weakly convex, almost flat along entire length; lateral impression very broad and shallow, obsolete. Pronotal disc slightly rugose, unevenly covered with mixed hollow-shaped moderate (0.03 mm) and fine (0.02 mm) punctures. Lateral impression with numerous large (0.08 mm) but very shallow punctures with wrinkled intervals. Large punctures along outer margin of lateral callus absent. Pronotum entirely marginate and ciliate anteriorly, marginate posteriorly. Anterior and posterior setiferous pores of pronotum absent.

Prothoracic hypomeron weakly convex, with weak impression and obsolete wrinkles laterally. Anterior cavity and basal fold of prothoracic hypomeron absent.

Metasternum entirely marginate anteriorly.

Elytron with obsolete humeral callus. Elytral punctures moderate to large (0.03–0.06 mm), hollow-shaped, arranged in regular, slightly paired rows, more or less visible among punctures in intervals. Interspaces between punctures slightly convex, rugose, covered by fine to moderate (0.02–0.03 mm) punctures. 5^th row with 15–18 punctures. Scutellar row with 5 punctures. Sutural furrow not very long, distinct at apical slope. Elytral epipleuron inclined outside, visible along entire length in lateral view, with sparse setae apically.

Hind wing absent.

1^st–3^rd tarsomeres with entire sole in both sexes, narrow in female, broadened in male. Male 1^st fore-tarsomere 0.9 X as wide as long, 0.8 X as wide as 3^rd tarsomere, 2.3 X as wide as claw tarsomere.

Pygidium with shallow longitudinal impression in basal ½ only in both sexes.

Last abdominal ventrite simple, convex in both sexes, with apical margin emarginate in male, twice S-shaped in female.

Aedeagus (Figs 4B, 8D) tube-shaped, with dorsal side concave along most of its length, 7 X as long as wide, with parallel lateral sides at most of its length, rectilinearly narrowed at sides of apical orifice, and bearing narrow apical projection. Apical projection rounded at the top. Aedeagus without apical denticles. Aedeagus strongly curved in lateral view, with apical part recurved upward. Flagellum tube-shaped, simple, exposed. Apical orifice 0.2 X as long as aedeagus length, cover plate 0.2 X as long as apical orifice. Pores present on lateral sides of apical orifice (in lateral view).

Female spermatheca present, but rudimentary, whip-shaped.

Body length: 7.7–7.9 mm (male), 8.4 mm (female); aedeagus 3.2 mm long, 0.5 mm wide.

Comments. After the original description (Daccordi et al., 2011), male last abdominal ventrite is without emargination. I believe, it is incorrect. In the available male (paratype), last abdominal ventrite has deeply emarginate apical margin.

Chrysolina yunnana Lopatin, 2008: 836

Material examined: Holotype: China, Yunnan Province, WNW of Zhongdian Town, 4,100–4,300 m, 27°53′53″N / 99°32′27″E; 27°54′01″N / 99°32′14″E, 14.07.2007, I. Belousov, I. Kabak leg.: male (ZIN); paratype: China, Yunnan Province, WNW of Zhongdian Town, 4,000 m, 27°54′03″N / 99°32′45″E, 13.07.2007, I. Belousov, I. Kabak leg.: 1 female (ZIN).

Description. Body elongate oval, 1.7 X as long as wide (Fig. 10B). Dorsum dark violet with green reflection on elytra or dark violet mixed with green, ventral side and legs dark green, with underside of 1^st and 2^nd antennomeres rufous. Dorsum entirely sericeous, microreticulate and micropunctulate.

Supraorbital line deep, long, but not reaching antennal insertion. Base of antenna 2.2 X closer to clypeus than to eye. Last maxillary palpomere narrow, barrel-shaped, obliquely truncate apically, 1.8 X as long as wide, 1.4 X as long as penultimate palpomere, 1.1 X as wide as the latter, similar in both sexes. Antenna 0.4 X as long as body. 7^th antennomere 1.5 X as long as wide, 9^th antennomere 1.4 X as long as wide.

Pronotum 2.0 X as wide as long, broadest basally or just before base, with lateral sides slightly rounded in posterior ½ and more rounded in anterior ½, more convergent forward than backward (when it broadest before base), with anterior side deeply emarginate, anterior angles rounded. Lateral callus weakly convex along entire length; lateral impression very broad, shallow. Pronotal disc evenly covered with fine to moderate (0.02–0.03 mm) punctures with some larger punctures basally and laterally. Lateral impression with numerous (about 30) large (0.06 mm) punctures. Moderate (0.03 mm) punctures present along outer margin of lateral callus. Pronotum entirely marginate and ciliate anteriorly, marginate posteriorly. Anterior and posterior setiferous pores of pronotum absent.

Prothoracic hypomeron almost flat, very weakly convex, with obsolete impression covered by obsolete wrinkles laterally. Anterior cavity of hypomeron small, basal fold absent.

Metasternum entirely marginate anteriorly.

Elytron without humeral callus. Elytral punctures moderate (0.03 mm), hollow-shaped, arranged in regular, paired rows. Interspaces between punctures slightly convex, rugose, covered by fine (0.02 mm) punctures and fine sparse wrinkles. 5^th row with 13–16 punctures. Scutellar row with 2 punctures. Sutural furrow very long, distinct at entire length of apical slope. Elytral epipleuron inclined outside, visible along entire length in lateral view, with only few setae apically.

Hind wing absent.

1^st–3^rd tarsomeres with entire sole in both sexes, narrow in female, very broad in male. Male 1^st fore-tarsomere 1.1 X as wide as long, 1.1 X as wide as 3^rd tarsomere, 3.7 X as wide as claw tarsomere.

Pygidium with shallow longitudinal impression in basal ½–⅔ only in both sexes.

Last abdominal ventrite simple, convex, with apical margin broadly truncate and bearing marginal furrow in both sexes.

Aedeagus (Figs 7B, 8E) strongly flattened dorso-ventrally, 3.1 X as long as wide, slightly broadened toward apical orifice, then roundly narrowed at sides of apical orifice transversely truncate at apex. Aedeagus without apical denticles. Aedeagus strongly curved, arc-shaped in lateral view. Flagellum narrow, simple, exposed. Apical orifice 0.3 X as long as aedeagus length, cover plate 0.5 X as long as apical orifice, with deep triangular emargination. Pores present on lateral sides apically.

Female spermatheca absent.

Body length: 8.7 mm (male), 7.6–8.5 mm (female); aedeagus 2.5 mm long, 0.8 mm wide.

Description of new species

Material examined: Holotype: China, Yunnan Province, Deqen, Tuoxia Highway, Mt. Range between Xiaruolisuzuxiang & Yezhizhen, 4,250 m, 27°42′50″ N / 99°11′27″ E, 11.6.2013, I. Belousov, I. Kabak, G. Davidian leg.: male (ZIN); paratype: with the same label: 1 female (BC).

Description. Male (holotype) (Fig. 10C). Body broadly oval, 1.5 X as long as wide. Dorsum green with diffuse violet pattern on pronotum and narrow violet elytral sutural and lateral stripes, ventral side and legs dark green mixed with violet, antennae black with underside of 1^st and 2^nd antennomeres rufous. Dorsum moderately shining, head and elytra microreticulate and micropunctulate, pronotum smooth.

Supraorbital line wide, shallow, short. Base of antenna 3 X closer to clypeus than to eye. Last maxillary palpomere narrow, barrel-shaped, obliquely truncate apically, 1.3 X as long as wide, similar to penultimate palpomere in length and width. Antenna 0.5 X as long as body. 7^th antennomere 1.7 X as long as wide, 9^th antennomere 1.6 X as long as wide.

Pronotum 1.9 X as wide as long, broadest basally, with lateral sides almost straight in posterior ½ and slightly rounded in anterior ½, convergent forward, with anterior side deeply emarginate, anterior angles roundly triangular. Lateral callus weakly convex along entire length; lateral impression broad, shallow, slightly deepened near base. Pronotal disc evenly covered with fine (0.02 mm) punctures. Lateral impression with not numerous (about 10) large (0.05–0.08 mm), partly coalescent, shallow punctures. Moderate (0.03 mm) punctures present along outer margin of lateral callus. Pronotum entirely marginate and ciliate anteriorly, entirely marginate posteriorly. Anterior and posterior setiferous pores of pronotum absent.

Prothoracic hypomeron weakly convex, with shallow impression covered by obsolete wrinkles laterally. Anterior cavity and basal fold absent.

Metasternum entirely marginate anteriorly.

Elytron without humeral callus. Elytral punctures moderate to large (0.03–0.06 mm), hollow-shaped, arranged in regular, paired rows. Interspaces between punctures slightly convex, rugose, covered by fine (0.02 mm) punctures and fine sparse wrinkles. 5^th row with 10 punctures. Scutellar row with 4 punctures. Sutural furrow very long, distinct along entire length of apical slope. Elytral epipleuron inclined outside, visible along entire length in lateral view, with sparse setae apically.

Hind wing absent.

1^st–3^rd tarsomeres with entire sole, moderately broadened. 1^st fore-tarsomere as wide as long, 1.1 X as wide as 3^rd tarsomere, 2.8 X as wide as claw tarsomere.

Pygidium with shallow longitudinal impression in basal ½ only.

Last abdominal ventrite simple, convex, with apical margin shallowly emarginate and bearing marginal furrow.

Aedeagus (Figs 4D, 8F) strongly flattened, 3 X as long as wide, slightly broadened toward apical orifice, then roundly narrowed at sides and obtusely 3-lobed at apex, with 2 large semi-transparent areas near apical margin. Aedeagus without apical denticles. Aedeagus strongly curved, arc-shaped in lateral view, with apical part slightly curved upward. Flagellum narrow, simple, exposed. Apical orifice 0.3 X as long as aedeagus length, cover plate 0.3 X as long as apical orifice, without triangular emargination. Fine pores present on dorsal side apically.

Body length 7.4 mm; aedeagus 2.1 mm long, 0.7 mm wide.

Description of female (paratype). Body 8.2 mm long; maxillary palpi similar to those in male; pronotum micropunctulate; 1^st–3^rd tarsomeres narrow, bearing entire sole; last abdominal ventrite broadly truncate; spermatheca absent.

Differential diagnosis. The new species is morphologically close to Ch. yunnana and differs in aedeagus bearing 2 semi-transparent areas apically, cover plate without triangular emargination, apical part slightly curved upward. The new species externally resembles the members of the subgenus Chrysolina (Volosatik) and differs in aedeagus apically broad, flattened, broadly truncate, and in pronotal lateral impression filled with large punctures.

Material examined: Holotype: China, Yunnan Province, Deqen, Tuoxia Highway Mt. Range between Xiaruolisuzuxiang & Yezhizhen, 4,250 m, 27°42′11″ N / 99°11′31″ E, 12.6.2013, I. Belousov, I. Kabak, G. Davidian leg.: male (ZIN); paratypes: with the same label: 5 males (BC); China, Yunnan Province, Deqen, Tuoxia Highway Mt. Range between Xiaruolisuzuxiang & Yezhizhen, 4,080 m, 27°40′2″ N / 99°10′29″ E, 13.6.2013, I. Belousov, I. Kabak, G. Davidian leg.: 1 male (BC); China, Yunnan Province, Deqen, Tuoxia Highway Mt. Range between Xiaruolisuzuxiang & Yezhizhen, 4,025 m, 27°42′50″ N / 99°11′27″ E, 11.6.2013, I. Belousov, I. Kabak, G. Davidian leg.: 1 male, 1 female (ZIN), 3 males, 2 females (BC); China, Yunnan Province, Mekong, Yangtze div., Baihaizi Lake vicinity, 3915 m, 27°49′39″ N / 99°07′17″ E, 5.6.2017, I. Belousov, G. Davidian, I. Kabak leg.: 1 male, 1 female (ZIN), 1 male, 1 female (BC); China, Yunnan Province, Mekong, Yangtze div., 4.8 km ENE of Wadang, 3,965 m, 27°49′09″ N / 99°07′51″ E, 4.6.2017, I. Belousov, G. Davidian, I. Kabak leg.: 1 male (BC).

Description. Male (holotype) (Fig. 10D). Body elongate oval, 1.7 X as long as wide. Dorsal side dark bronze; underside and legs black with weak metallic reflection; base of trochanters reddish; antennae piceous with 1^st and 2^nd antennomeres rufous below. Dorsum moderately shining, microreticulate and micropunctulate.

Supraorbital line distinct above eye. Base of antenna 2.0 X closer to clypeus than to eye. Last maxillary palpomere narrow, barrel-shaped, obliquely truncate apically, 1.5 X as long as wide, 1.1 X as long as penultimate palpomere, as wide as the latter. Antenna 0.5 X as long as body. 7^th antennomere 1.3 X as long as wide, 9^th antennomere 1.5 X as long as wide.

Pronotum 1.8 X as wide as long, broadest at mid-length, with lateral sides rounded, more convergent forward than backward, with anterior side moderately emarginate, anterior angles rounded. Lateral callus weakly convex along entire length; lateral impression broad, weak, forming two weak cavities, first at base and second in anterior ½. Pronotal disc evenly covered with dense, fine (0.02 mm) punctures. Lateral impression with several (about 10) moderate (0.03 mm) punctures. Four moderate punctures present along outer margin of lateral callus. Pronotum entirely marginate and ciliate anteriorly, entirely marginate posteriorly. Anterior and posterior setiferous pores of pronotum absent.

Prothoracic hypomeron flat and covered with obsolete irregular wrinkles laterally. Anterior cavity and basal fold of prothoracic hypomeron absent.

Metasternum entirely marginate anteriorly.

Elytron with obsolete humeral callus. Elytral punctures moderate (0.03 mm), arranged in regular, equidistant rows, with intervals slightly convex, covered by very fine wrinkles. 5^th row with 24 punctures. Scutellar row with 3–7 punctures. Sutural furrow distinct at apical slope. Elytral epipleuron inclined outside, visible along entire length in lateral view, sparsely ciliate apically.

Hind wing absent.

1^st–3^rd tarsomeres very broad, with entire sole. 1^st fore-tarsomere 1.1 X as wide as long, as wide as 3^rd tarsomere, 3.2 X as wide as claw tarsomere.

Pygidium with shallow, obsolete longitudinal impression in basal ½ only.

Last abdominal ventrite simple, convex, with apical margin broadly truncate and bearing furrow.

Aedeagus (Figs 1I, 8G) flattened dorso-ventrally, 3 X as long as wide, with almost parallel lateral sides, hardly broadened toward apex, with apical margin transversely truncate, slightly rounded and bearing shallow emargination. Aedeagus with very fine apical denticles. Aedeagus arc-shaped, rather curved in lateral view. Flagellum narrow, simple, exposed. Apical orifice 0.3 X as long as aedeagus length, cover plate 0.7 X as long as apical orifice. Pores present along apical margin (viewed dorsally).

Body length 6.4 mm; aedeagus 1.8 mm long, 0.6 mm wide.

Description of female (paratypes). Body sericeous dorsally; 1^st–3^rd tarsomeres narrow, with entire sole; last abdominal ventrite simple, convex, with apical margin rounded; spermartheca absent.

Variability. Males 6.0–6.8 mm long, females 6.5–7.4 mm long. Dorsum rarely dark green with golden reflection. Pronotum broadest basally with lateral sides almost straight in basal ⅓ in some paratypes. Pronotal lateral impression from obsolete to moderate and developed along entire length. Elytral intervals from almost flat to slightly keel-shaped.

Differential diagnosis. The new species is externally very close to Ch. luobinwangi and differs in male aedeagus apically truncate but not triangular, with apical orifice covered by plate over most of its length.

Material examined: Holotype: China, Yunnan Province, Yulongshan mts., 3,300–3,500 m, 27°7′ N / 100°13′ E, 20–21.6.1993, V. Kubáň leg.: 1 male (ZIN); paratypes: China, Yunnan Province, 20 km N of Lijiang Yulongshan, 3,406 m, dry river, 27°5′39″ N / 100°13′45″ E, 30.5.2002, A. Konstantinov, M. Volkovitsh leg.: 2 males (IZCAS); China, Yunnan Province, 20 km N of Lijiang Yulongshan, 3,406 m, moss sifting, 27°5′39″ N / 100°13′45″ E, 30.5.2002, A. Konstantinov, M. Volkovitsh leg.: 1 female (ZIN); China, Yunnan Province, environs of Lijang lower Yak meadow, 2,800 m, 27°8′48″ N / 100°15′02″ E, 27.5.2002, A. Konstantinov, M. Volkovitsh leg.: 1 male, 1 female (USNM), 1 male (ZIN); China, N Yunnan Province, 30 km N of Lijiang, 3,000 m, 3.7.1990, L. and M. Bocák leg.: 1 male (ZIN), 1 male (BC); China, Yunnan Province, Yulongshan, 3,300 m, 27°7′ N / 100°14′ E, 9.7.1995, Bolm leg.: 1 male (BC); China, N Yunnan Province, environs of Heishui, 24–28.5.1993, R. Cervenka leg.: 2 females (DC), 1 female (BC); China, Yunnan Province, Yulongshan mts., 3,400 m, 27°7′ N / 100°13′ E, 20–21.6.1993, Bolm leg.: 1 female (ZIN); China, Yunnan Province, Yulongshan mts., Ganhaizi pass, 3,000–3,500 m, 27°6′ N / 100°15′ E, 18–23.7.1990, V. Kubáň leg.: 1 female (BC); China, Yunnan Province, Lijiang, 05.1995, S.J.L. Remond leg.: 1 male (BC); China, Yunnan Province, W Lijiang, W Yangtze, W Xinhuacun, 0.7 km NW of Daqingtou, 2,720 m, 26°56′23″ N / 99°52′16″ E, 1.6.2018, I. Belousov, I. Kabak leg.: 1 male, 3 females (BC); China, Yunnan Province, Hutiao gorge, Jinsha river, 2,000 m, 27.15 N / 100.09 E, 18–22.7.1992, D. Král leg.: 1 female (BC); China, Yunnan Province, Hutiao gorge, Jinsha river, 2,000 m, 27.15 N / 100.09 E, 18–22.7.1992, V. Kubáň leg.: 2 males (BC); China, Yunnan Province, Heishui, 35 km N of Lijiang, 27.13 N / 100.19 E, 1–19.7.1992, S. Bečvar leg.: 4 males, 4 females (DC), 1 female (ZIN).

Description. Male (holotype) (Fig. 14B). Body oval, 1.5 X as long as wide. Dorsal side dark bronze, ventral side and legs black with dark bronze reflection; antennae black with 1^st antennomere reddish below, 2^nd antennomere reddish. Head, pronotum, and elytra sericeous shining. Dorsum microreticulate, besides that pronotum micropunctulate.

Supraorbital line present above eye. Base of antenna 3 X closer to clypeus than to eye. Last maxillary palpomere narrow, barrel-shaped, obliquely truncate at apex, 1.4 X as long as wide, 1.5 X as long as penultimate palpomere and 1.1 X as wide as the latter. Antenna 0.5 X as long as body. 7^th antennomere 1.4 X as long as wide, 9^th antennomere 1.6 X as long as wide.

Pronotum 1.8 X as wide as long, broadest basally, with lateral sides rounded, convergent forward and obsoletely emarginate before base, with anterior side moderately emarginate, anterior angles roundly triangular. Lateral callus moderately convex along entire length; lateral impression broad, weak, developed along entire length. Pronotal disc evenly covered with dense, fine (0.02 mm) punctures. Lateral impression with about 10 large (0.05 mm) punctures in anterior ½ and near base. 4 or 5 moderate (0.03 mm) punctures present along outer margin of lateral callus. Pronotum entirely marginate and ciliate anteriorly, entirely marginate posteriorly. Anterior and posterior setiferous pores of pronotum absent.

Prothoracic hypomeron weakly convex, without lateral impression, with obsolete wrinkles laterally. Anterior cavity of prothoracic hypomeron small, basal fold absent.

Metasternum entirely marginate anteriorly.

Elytron without humeral callus. Elytral punctures moderate (0.03 mm), arranged in regular, paired (2–3, 4–5, 6–7, 8–9) rows, with intervals flat. Intervals with numerous fine (0.02 mm) punctures and sparse obsolete wrinkles. 5^th row with 24 punctures. Scutellar row consists of 4 punctures. Sutural furrow distinct at apical slope. Elytral epipleuron inclined outside, visible along entire length in lateral view, with sparse setae apically.

Hind wing absent.

1^st–3^rd tarsomeres narrow, with entire sole. 1^st fore-tarsomere 0.9 X as wide as long, 0.9 X as wide as 3^rd tarsomere, 2.3 X as wide as claw tarsomere.

Pygidium with shallow, obsolete longitudinal impression in basal ½ only.

Last abdominal ventrite simple, convex, with apical margin broadly emarginate and bearing marginal furrow.

Aedeagus (Figs 3A, 15D) strongly flattened dorso-ventrally, with dorsal side longitudinally concave at mid-length, 4 X as long as wide, almost parallel-sided, slightly narrowed at sides of apical orifice, with apical margin emarginate between 2 rounded lobes. Aedeagus without apical denticle. Aedeagus strongly curved near mid-length in lateral view. Flagellum simple, narrow. Apical orifice 0.3 X as long as aedeagus length, cover plate 0.1 X as long as apical orifice. Pores absent.

Body length 6.2 mm; aedeagus 2.4 mm long, 0.6 mm wide.

Description of female (paratypes). 1^st–3^rd tarsomeres narrow, with entire sole; last abdominal ventrite convex, with apical margin rounded; spermatheca present.

Variability. Males 5.6–8.4 mm long. Females 7.2–8.7 mm long. Pronotum sometimes broadest at mid-length.

Differential diagnosis. The new species resembles Ch. luluni sp. nov. and differs in elytral puncture rows regular, distinct, and in the aedeagus longer, 4 X as long as wide.

Comments. Female is ovoviviparous. Embryos with developed ocelli, mandibulae, setae on body segments, and legs were found inside the abdomen.

This species has a significant variability in the aedeagus shape (Fig. 3A–C). However: 1) no clear differences in the structure of the aedeagus were found, 2) the limits of variability in the shape of the aedeagus are less than between different species in the present subgenus, 3) all 13 known localities of this species come from the same, very limited area (Fig. 6), 4) externally all specimens are quite similar. All of the above allows me to assume that the available specimens belong to the same species.

Material examined: Holotype: China, Yunnan Province, N of Weixi, left tributary of Luozhua, 7.75 km E of Nianjua Mt., 4,080 m, 27°44′9″ N / 98°57′3″ E, 26.5.2016, I. Kabak, G. Davidian leg.: male (ZIN); paratypes: China, Yunnan Province, N of Weixi, 8.65 km W Laoantong, 7.35 km E of Nianjua Mt., 3,950 m, 27°44′41″ N / 98°56′58″ E, 16.6.2015, I. Belousov, I. Kabak, G. Davidian leg.: 1 male (BC); China, Yunnan Province, N of Weixi, left tributary of Luozhua, 7.75 km E of Nianjua Mt., 4,080 m, 27°44′9″ N / 98°57′3″ E, 15.6.2015, I. Belousov, I. Kabak, G. Davidian leg.: 1 male (BC).

Description. Male (holotype) (Fig. 16B). Body elongate oval, 1.7 X as long as wide. Dorsal side dark coppery, with pronotum, elytral suture and lateral margin dark green, underside and legs black; antennae piceous with 1^st and 2^nd antennomeres rufous below. Dorsum sericeous, micropunctulate and microreticulate.

Supraorbital line broad, developed above eye. Base of antenna 3.0 X closer to clypeus than to eye. Last maxillary palpomere narrow, barrel-shaped, 1.7 X as long as wide, 1.6 X as long as penultimate palpomere, 1.1 X as wide as the latter. Antenna 0.4 X as long as body. 7^th antennomere 1.1 X as long as wide, 9^th antennomere 1.2 X as long as wide.

Pronotum 2.0 X as wide as long, broadest at mid-length, with lateral sides rounded, more convergent forward than backward, with anterior side moderately emarginate, anterior angles rounded. Lateral callus moderately convex along entire length; lateral impression broad, moderate along entire length, slightly deepened and bearing trace of outer border basally. Pronotal disc evenly covered with dense, fine (0.02 mm) punctures. Lateral impression with not numerous (about 10) large (0.05–0.08 mm) punctures. Several moderate (0.03 mm) punctures present along outer margin of lateral callus. Pronotum entirely marginate and ciliate anteriorly, entirely marginate posteriorly. Anterior and posterior setiferous pores of pronotum absent.

Prothoracic hypomeron flat and covered with weak irregular wrinkles laterally. Anterior cavity small, basal fold of prothoracic hypomeron absent.

Metasternum entirely marginate anteriorly.

Elytron without humeral callus. Elytral punctures moderate (about 0.03 mm), arranged in equidistant undulate and partly irregular rows, 6^th and 7^th rows mostly irregular apically, with intervals weakly convex, covered by fine wrinkles. 5^th row with 35 punctures. Scutellar row with 2 punctures. Sutural furrow distinct at apical slope. Elytral epipleuron inclined outside, visible along entire length in lateral view, sparsely ciliate apically.

Hind wing absent.

1^st–3^rd tarsomeres moderately broadened, with entire sole. 1^st fore-tarsomere as wide as long, 0.7 X as wide as 3^rd tarsomere, 2.5 X as wide as claw tarsomere.

Pygidium with weak, shallow longitudinal impression in basal ½ only.

Last abdominal ventrite simple, convex, with apical margin broadly truncate, with furrow along apical margin.

Aedeagus (Figs 3F, 8I) strongly flattened dorso-ventrally, 3.3 X as long as wide, with almost parallel lateral sides, hardly broadened toward apex, apically broadly trapeziform with lateral sides slightly rounded, and apical margin slightly emarginate. Aedeagus without distinct apical denticles, but with two small swellings at apical margin ventrally. Aedeagus moderately curved in lateral view. Flagellum tube-shaped, simple, exposed. Apical orifice 0.3 X as long as aedeagus length, cover plate 0.3 X as long as apical orifice. Pores present near apical margin dorsally and ventrally.

Body length 6.8 mm; aedeagus 2.0 mm long, 0.6 mm wide.

Variability. Body length 7.0–7.1 mm. In some specimens, dorsum mostly dark green with dark coppery spots on head and pronotum and elytral intervals 1–3 or 1–5.

Differential diagnosis. The new species is externally some similar with Ch. luobinwangi and Ch. selene sp. nov. and differs in dorsum two-colored (coppery with green) but not entirely bronze, in male 1^st–3^rd fore-tarsomeres less broadened, aedeagus apically roundly truncate but not triangular, with flagellum broader, tube-shaped, and in the pronotal lateral impression more developed.

Comments. Female is unknown.

Material examined: Holotype: China, Yunnan Province, Yulongshan mts., 4,000 m, 27.02 N / 100.11 E, 27.5.1993, Bolm leg.: male (ZIN); paratypes: China, Yunnan Province, Yulongshan mts., 3,600–4,100 m, 27.02 N / 100.11 E, 27.5.1993, V. Kubáň leg.: 1 male (BC), 1 male (DC); China, NW Yunnan Province, Yulongxue Shan, W slope, between Bahanchang and Tuguancun, meadow / mixed forest margin, 2,700 m, 27.03 N / 100.08 E, 2.7.1996, B. Březina leg.: 1 male (DC).

Description. Male (holotype) (Fig. 14C). Body broadly oval, 1.5 X as long as wide. Dorsal side dark bronze; underside and legs black with bronze reflection; antennae black with 1^st and 2^nd antennomeres reddish below. Dorsum sericeous, microreticulate, besides that head and pronotum micropunctulate.

Supraorbital line absent. Base of antenna 2.0 X closer to clypeus than to eye. Last maxillary palpomere narrow, barrel-shaped, obliquely truncate at apex, 1.4 X as long as wide, 1.2 X as long as penultimate palpomere and 1.1 X as wide as the latter. Antenna 0.6 X as long as body. 7^th antennomere 1.3 X as long as wide, 9^th antennomere 1.2 X as long as wide.

Pronotum 1.9 X as wide as long, broadest at mid-length, with lateral sides rounded, more convergent forward than backward, with anterior side moderately emarginate, anterior angles rounded. Lateral callus weakly convex along entire length; lateral impression almost absent. Pronotal disc evenly covered with dense, fine (0.02 mm) punctures. Lateral sides (at the place of impression) with only few (4–7) large (0.05 mm) punctures near base. Moderate (0.03 mm) punctures present along outer margin of lateral callus. Pronotum entirely marginate and ciliate anteriorly, entirely marginate posteriorly. Anterior and posterior setiferous pores of pronotum absent.

Prothoracic hypomeron convex, with weak lateral impression covered by obsolete wrinkles. Anterior cavity of prothoracic hypomeron small, basal fold absent.

Metasternum entirely marginate anteriorly.

Elytron without humeral callus. Elytral punctures moderate (0.03 mm), partly arranged in irregular, slightly paired (2–3, 4–5, 6–7, 8–9) rows, with intervals flat. Rows hardly visible because of numerous fine (0.02 mm) punctures in intervals. Intervals with sparse obsolete wrinkles. 5^th row with 25 punctures. Scutellar row with 10 punctures. Sutural furrow distinct at apical slope. Elytral epipleuron inclined outside, visible along entire length in lateral view, with only few setae apically.

Hind wing absent.

1^st–3^rd tarsomeres slightly broadened, with entire sole. 1^st fore-tarsomere as wide as long, 0.8 X as wide as 3^rd, 2.3 X as wide as claw tarsomere.

Pygidium with shallow, obsolete longitudinal impression in basal ½ only.

Last abdominal ventrite simple, convex, with apical margin broadly truncate.

Aedeagus (Figs 1K, 15E) strongly flattened dorso-ventrally, 2.7 X as long as wide, broadened from its base toward the base of apical orifice, and then rectilinearly narrowed toward apex, with apical margin broadly truncate and slightly emarginate. Aedeagus without apical denticles, but with 2 small swellings at apical margin ventrally. Aedeagus strongly curved near mid-length in lateral view. Flagellum simple, narrow. Apical orifice 0.3 X as long as aedeagus length, cover plate 0.5 X as long as apical orifice. Pores present on ventral side of aedeagus in apical ½ and on the sides of apical orifice.

Body length 5.8 mm; aedeagus 1.6 mm long, 0.6 mm wide.

Variability. Male 5.4 mm long. Pronotal disc with greenish tint.

Differential diagnosis. The new species resembles Ch. shuyongi Ge et Daccordi in: Ge et al., 2011 and differs in male body broader (1.5 X as long as wide, while 1.9 X as long as wide in Ch. shuyongi), elytral puncture rows irregular, hardly traced, while rows regular in Ch. shuyongi, and in the aedeagus narrowed (but not roundly broadened) at sides of apical orifice, and not recurved dorsally at apex (viewed laterally). It differs also from Ch. konstantinovi sp. nov. in elytral rows irregular, and aedeagus shorter, 2.7 X as long as wide.

Comments. Female is unknown.

Material examined: Holotype: China, NW Sichuan Province, 16.8 km SSW Ganzi, 4,490 m, 31°28′18″ N / 99°56′21″ E, 12.7.2016, I. Belousov, I. Kabak leg.: male (ZIN); paratypes: with the same label: 1 male, 1 female (ZIN), 1 male (BC); China, W Sichuan Province, pass 17 km SE-SSE of Ronggai, alpine meadows, screes, 4,800 m, 30°59′ N / 99°00′ E, 15–18.6.2018, J. Kaláb leg.: 1 female (BC); China, NW Sichuan Province, 24 km S Ganzi, 4,735 m, 31°24′1″ N / 99°58′42″ E, 14.7.2016, I. Belousov, I. Kabak leg.: 1 male (BC); China, Sichuan Province, Kawaluri Mt. environs, 4,415 m, 31°28′30″ N / 100°14′24″ E, 8.7.2016, I. Belousov, I. Kabak leg.: 1 male (RC).

Description. Male (holotype) (Fig. 16C). Body broadly oval, 1.5 X as long as wide. Dorsal side dark bronze; underside and legs black with bronze reflection; antennae piceous with 1^st and 2^nd antennomeres rufous below. Dorsum moderately shining with sericeous reticulated spots on elytra, with pronotum distinctly microreticulate and micropunctulate, elytra with strongly reticulated spots and obsolete reticulation in interspaces (Fig. 2C).

Supraorbital line distinct above eye. Base of antenna 1.8 X closer to clypeus than to eye. Last maxillary palpomere narrow, barrel-shaped, obliquely truncate apically, 1.3 X as long as wide, 1.2 X as long as penultimate palpomere, 1.1 X as wide as the latter. Antenna 0.4 X as long as body. 7^th antennomere as long as wide, 9^th antennomere 1.1 X as long as wide.

Pronotum 1.9 X as wide as long, broadest before base, with lateral sides rounded, more convergent toward apex than toward base, with anterior side moderately emarginate, anterior angles rounded. Lateral callus weakly convex along entire length; lateral impression broad, obsolete along most part of its length, slightly more developed only at base. Pronotal disc evenly covered with dense, fine to moderate (0.02–0.03 mm) punctures. Lateral impression with several large (0.05–0.06 mm), partly coalescent punctures only at base. Large punctures along outer margin of lateral callus absent. Pronotum entirely marginate and ciliate anteriorly, entirely marginate posteriorly. Anterior and posterior setiferous pores of pronotum absent.

Prothoracic hypomeron weakly convex, without lateral impression, covered with weak transverse wrinkles laterally. Anterior cavity and basal fold of prothoracic hypomeron absent.

Metasternum entirely marginate anteriorly.

Elytron without humeral callus. Elytral punctures fine (0.02 mm), arranged in regular, equidistant or slightly paired rows, with intervals very weakly convex, covered by irregular fine punctures and fine wrinkles. Each puncture in rows surrounded by large (0.2 mm) spot of strong reticulation (Fig. 2C). 5^th row with 22 punctures. Scutellar row with 2 punctures. Sutural furrow short (0.8 mm), developed at posterior part of apical slope. Elytral epipleuron inclined outside, visible along entire length in lateral view, sparsely ciliate apically.

Hind wing absent.

1^st–3^rd tarsomeres moderately broadened, with entire sole. 1^st fore-tarsomere as wide as long, 0.9 X as wide as 3^rd tarsomere, 2.9 X as wide as claw tarsomere.

Pygidium with weak, shallow longitudinal impression in basal ½ only.

Last abdominal ventrite simple, convex, with apical margin broadly truncate.

Aedeagus (Figs 1J, 15A) flattened dorso-ventrally, 3.1 X as long as wide, with parallel lateral sides, with apical margin rounded. Aedeagus with large, obtuse, lobe-shaped apical denticles on ventral side. Aedeagus strongly curved in lateral view. Flagellum narrow, simple, exposed. Apical orifice 0.3 X as long as aedeagus length, cover plate 0.3 X as long as apical orifice. Pores present along lateral side of apical orifice (viewed laterally).

Body length 6.3 mm; aedeagus 2.5 mm long, 0.8 mm wide.

Description of female (paratypes). 1^st–3^rd tarsomeres narrow, with entire sole; last abdominal ventrite slightly swollen, with apical margin rounded; spermatheca absent.

Variability. Males 6.1–6.8 mm long, females 6.2–7.4 mm long. Male 7^th antennomere 1.0–1.1 X as long as wide; 1^st fore-tarsomere 2.5–2.9 X as wide as claw tarsomere; 5^th elytral row with 16–22 punctures.

Differential diagnosis. The new species externally differs from all other members of the present subgenus by the elytral relief: rows of strongly reticulated rounded spots. Similar elytral relief also presents in Ch. melchiori Bieńkowski, 2023, however, the elytral punctures are entirely irregular in the last species (Bieńkowski 2023).

Comments. Female is ovoviviparous. Embryos with developed ocelli, mandibulae, setae on body segments, and legs were found inside the abdomen.

Material examined: Holotype: China, Sichuan Province, S of Muli, 3,975–4,120 m, 27°43′16″ N / 101°15′13″ E − 27°43′03″ N / 101°15′05″ E, 22.5.2012, I. Belousov, G. Davidian, I. Kabak, A. Korolev leg.: male (ZIN); paratypes: with the same label: 1 male (ZIN), 2 males (BC).

Description. Male (holotype) (Fig. 14A). Body elongate oval, 1.7 X as long as wide. Dorsal side dark bronze; underside and legs black with bluish and greenish metallic reflection; antennae black with 1^st and 2^nd antennomeres reddish below. Dorsum moderately shining, micropunctulate and obsoletely microreticulate.

Supraorbital line distinct above eye. Base of antenna 2.0 X closer to clypeus than to eye. Last maxillary palpomere narrow, barrel-shaped, 1.5 X as long as wide, 1.4 X as long as penultimate palpomere and 1.1 X as wide as the latter. Antenna 0.4 X as long as body. 7^th antennomere 1.1 X as long as wide, 9^th antennomere as long as wide.

Pronotum 1.9 X as wide as long, broadest at mid-length, with lateral sides rounded, more convergent forward than backward, with anterior side weakly emarginate, anterior angles rounded. Lateral callus weakly convex along entire length; lateral impression broad, obsolete. Pronotal disc evenly covered with dense, fine (0.02 mm) punctures. Lateral impression without punctures larger than those on disc. Large punctures along outer margin of lateral callus absent. Pronotum entirely marginate and ciliate anteriorly, entirely marginate posteriorly. Anterior and posterior setiferous pores of pronotum absent.

Prothoracic hypomeron convex, without lateral impression, with weak transverse wrinkles laterally. Anterior cavity of prothoracic hypomeron small, basal fold absent.

Metasternum entirely marginate anteriorly.

Elytron without humeral callus. Elytral punctures moderate (0.03 mm), arranged in regular, partly undulate, equidistant rows, with intervals flat, smooth, without fine wrinkles. 5^th row with 30 punctures. Scutellar row with 8 punctures. Sutural furrow distinct at apical slope. Elytral epipleuron inclined outside, visible along entire length in lateral view, sparsely ciliate apically.

Hind wing absent.

1^st–3^rd tarsomeres slightly broadened, with entire sole. 1^st fore-tarsomere as wide as long, as wide as 3^rd tarsomere, 2.6 X as wide as claw tarsomere.

Pygidium with shallow, obsolete longitudinal impression in basal ½ only.

Last abdominal ventrite simple, convex, with apical margin slightly emarginate and bearing furrow.

Aedeagus (Figs 3D, 15C) flattened dorso-ventrally, 4.2 X as long as wide, with almost parallel lateral sides, slightly narrowed toward apex, with apical margin roundly triangular. Aedeagus without apical denticles. Aedeagus arc-shaped, rather curved in lateral view. Flagellum absent. Apical orifice 0.6 X as long as aedeagus length, cover plate 0.6 X as long as apical orifice. Pores near apex absent.

Body length 5.8 mm; aedeagus 2.1 mm long, 0.5 mm wide.

Variability. Males 5.5–5.8 mm long. Pronotal lateral impression wide, shallow along entire length, slightly deepened basally.

Differential diagnosis. The new species externally close to Ch. hongyuanensis and Ch. pascolii sp. nov. and differs in prothoracic hypomeron convex, not flat, and aedeagus apically roundly triangular, neither anchor-shaped, nor right triangular.

Comments. Female is unknown.

Material examined: Holotype: China, W Sichuan Province, mts. 13 km W-WNW of Daolu, alpine meadows, screes, 4,600 m, 31°00′ N / 100°59′ E − 30°59′ N / 101°00′ E, 4–8.6.2014, J. Kaláb leg.: male (ZIN); paratypes: with the same label: 2 males, 1 female (BC), 3 males, 1 female (DC).

Description. Male (holotype) (Fig. 14D). Body oval, 1.6 X as long as wide. Dorsal side dark bronze; underside and legs black with weak bronze reflection; antennae piceous with 1^st and 2^nd antennomeres reddish below. Dorsum moderately shining, micropunctulate and microreticulate.

Supraorbital line distinct above eye. Base of antenna 2.5 X closer to clypeus than to eye. Last maxillary palpomere narrow, barrel-shaped, 1.3 X as long as wide, 1.3 X as long as penultimate palpomere and as wide as the latter. Antenna 0.5 X as long as body. 7^th antennomere 0.9 X as long as wide, 9^th antennomere 1.1 X as long as wide.

Pronotum 1.9 X as wide as long, broadest at base, with lateral sides weakly rounded, convergent forward, with anterior side weakly emarginate, anterior angles rounded. Lateral callus weakly convex along entire length; lateral impression broad, obsolete, slightly deepened in basal ½. Pronotal disc evenly covered with dense, fine (0.02 mm) punctures. Lateral impression with sparse (15–16) moderate (0.03 mm) punctures. Several moderate (0.03 mm) punctures present along outer margin of lateral callus. Pronotum entirely marginate and ciliate anteriorly, entirely marginate posteriorly. Anterior and posterior setiferous pores of pronotum absent.

Prothoracic hypomeron weakly convex, without lateral impression, with weak transverse wrinkles laterally. Anterior cavity of prothoracic hypomeron and basal fold absent.

Metasternum entirely marginate anteriorly.

Elytron without humeral callus. Elytral punctures from moderate to large (0.03–0.05 mm), arranged in partly irregular undulate paired rows. Rows 6^th and 7^th mostly irregular. Intervals flat, with dense fine (0.02 mm) punctures and fine wrinkles. 5^th row with 18 punctures. Scutellar row with 5–7 punctures. Sutural furrow distinct at apical slope. Elytral epipleuron inclined outside, visible along entire length in lateral view, with only few setae apically.

Hind wing absent.

1^st–3^rd tarsomeres slightly broadened, with entire sole. 1^st fore-tarsomere as wide as long, as wide as 3^rd tarsomere, 2.6 X as wide as claw tarsomere.

Pygidium with shallow, obsolete longitudinal impression in basal ½ only.

Last abdominal ventrite simple, convex, with apical margin straight and bearing furrow.

Aedeagus (Figs 7C, 15H) tube-shaped, with dorsal side concave at mid-length, 4.4 X as long as wide, with parallel lateral sides from base till apical orifice, with apical margin elongate triangular and narrowly truncate. Aedeagus with small apical denticles ventrally. Aedeagus arc-shaped, rather curved in basal ½ (viewed laterally). Flagellum narrow, simple, exposed. Apical orifice 0.3 X as long as aedeagus length, cover plate 0.2 X as long as apical orifice. Pores present at lateral sides at level of apical orifice (viewed laterally).

Body length 5.7 mm; aedeagus 2.2 mm long, 0.5 mm wide.

Description of female (paratype). Body 6.6 mm long, spermatheca present.

Variability. Males 5.7–6.0 mm long.

Differential diagnosis. The new species externally close to Ch. hongyuanensis and Ch. pakhomovae sp. nov. and differs in aedeagus triangular apically and bearing narrow flagellum. It also differs from Ch. romantsovi sp. nov. in aedeagus triangular apically, and pronotal lateral impression deepened basally.

Material examined: Holotype: China, N Sichuan Province, pass between Songpan town and Juzaigou, 3,700 m, 33°01′56″ N / 103°41′48″ E, 17.7.2005, V. Patrikeev leg.: male (ZIN); paratypes: China, E Qinghai Province, 33 km SW Sinine city, 3,137 m, 36°15′34″ N / 101°34′00″ E, V. Patrikeev leg.: 2 males (RC, BC) [locality of paratypes looks like incorrect, see Comments].

Description. Male (holotype) (Fig. 20D). Body elongate oval, 1.7 X as long as wide. Dorsal side dark bronze; ventral side and legs black; antennae piceous with 1^st antennomere below rufous, 2^nd mostly rufous. Head and pronotum sericeous, elytra moderately shining, dorsum micropunctulate and microreticulate.

Supraorbital line distinct above eye. Base of antenna 1.2 X closer to clypeus than to eye. Last maxillary palpomere narrow, barrel-shaped, 1.2 X as long as wide, similar with penultimate palpomere in length and width. Antenna 0.5 X as long as body. 7^th antennomere 1.5 X as long as wide, 9^th antennomere 1.2 X as long as wide.

Pronotum 2.0 X as wide as long, broadest at base, with lateral sides convergent forward, almost rectilinear in basal ½ and rounded in anterior ½, with anterior side moderately emarginate, anterior angles roundly triangular. Lateral callus weakly convex along entire length; lateral impression shallow along entire length. Pronotum covered with dense, fine (0.02 mm) punctures, lateral impressions with 7–10 moderate to large (0.03–0.05 mm) punctures. Several moderate punctures present along outer margin of lateral callus. Pronotum entirely marginate and ciliate anteriorly, entirely marginate posteriorly. Anterior and posterior setiferous pores of pronotum absent.

Prothoracic hypomeron weakly convex, without lateral impression, with obsolete wrinkles laterally. Anterior cavity of prothoracic hypomeron and basal fold absent.

Metasternum entirely marginate anteriorly.

Elytron with obsolete humeral callus. Elytral punctures moderate (0.03 mm), hollow-shaped, arranged in regular, equidistant, sparse rows. Intervals almost flat (obsoletely convex), with sparse fine (0.02 mm) punctures and sparse fine wrinkles. 5^th row with 10 punctures. Scutellar row with 1–2 punctures. Sutural furrow distinct in posterior part of apical slope. Elytral epipleuron inclined outside, visible along entire length in lateral view, with only few setae apically.

Hind wing absent.

1^st–3^rd tarsomeres slightly broadened, with entire sole. 1^st fore-tarsomere 0.9 X as wide as long, 0.9 X as wide as 3^rd tarsomere, 2.4 X as wide as claw tarsomere.

Pygidium with shallow, obsolete longitudinal impression in basal ½ only.

Last abdominal ventrite simple, convex, with apical margin broadly truncate and bearing furrow.

Aedeagus (Fig. 2A, B) slightly flattened dorso-ventrally, 4.5 X as long as wide, with parallel lateral sides along almost entire length, with apical margin rounded. Aedeagus with 2 small obtuse apical denticles on ventral side. Aedeagus arc-shaped, rather curved. Flagellum narrow, simple, exposed. Apical orifice 0.3 X as long as aedeagus length, cover plate 0.2 X as long as apical orifice. Pores present at lateral sides at level of apical orifice (viewed laterally).

Body length 5.2 mm; aedeagus 1.8 mm long, 0.4 mm wide.

Variability. 5^th elytral row with 12 punctures. Males 4.7 mm long.

Differential diagnosis. The new species externally close to Ch. romantsovi sp. nov. and differs in aedeagus rounded apically, pronotal lateral impression more distinct and bearing moderate to large punctures. It also resembles Ch. (Allohypericia) mauroi by the external characters and aedeagus structure and differs in distinct bronze color dorsally (dorsal side black with weak bronze reflection in Ch. mauroi), pronotal lateral impression evenly shallow along entire length, not deepened in basal 1/3 (impression with narrow furrow in basal 1/3 in Ch. mauroi), covered by sparse (7–10) separate punctures (18–20 partly coalescent punctures in Ch. mauroi), and in fore-tarsomeres 1^st–3^rd narrower.

Etymology. The new species is named after the Russian entomologist V.M. Patrikeev, who collected the type specimens.

Comments. Female is unknown.

The shape of the aedeagus in Ch. patrikeevi sp. nov. probably represents a generalized aedeagus plan. A similar shape of aedeagus also presents in such unrelated species as Ch. claripes Lopatin, 2002, Ch. jinxiaoae Ge in Daccordi et al. 2011, Ch. libaii Daccordi et Ge in Daccordi et al. 2011, and Ch. lijieae Daccordi et Yang in Daccordi et al. 2011; Bieńkowski 2019).

Material examined: Holotype: China, N Yunnan Province, mts. 15 km W of Zhongdian, 3,800–4,200 m, 27.49 N / 99.34 E, 22–24.6.1994, J. Farkač, D. Král leg.: male (ZIN); paratype: China, N Yunnan Province, Xue Shan near Zhongdian, 4,000–4,100 m, 27.49 N / 99.34 E, 23.6.1996, J. Farkač, P. Kabátek, A. Smetana leg.: 1 male (DC).

Description. Male (holotype) (Fig. 20B). Body broadly oval, 1.5 X as long as wide. Body dark bronze, with weak coppery reflection on head and pronotum, base of trochanters rufous, antennae piceous with 1^st–2^nd antennomeres rufous below. Dorsum moderately shining, microreticulate, besides that head and pronotum micropunctulate.

Supraorbital line deep, long, but not reaching antennal insertion. Base of antenna 2.5 X closer to clypeus than to eye. Last maxillary palpomere narrow, barrel-shaped, obliquely truncate apically, 1.4 X as long as wide, 1.4 X as long as penultimate palpomere, 1.1 X as wide as the latter. Antenna 0.5 X as long as body. 7^th antennomere 1.6 X as long as wide, 9^th antennomere 1.7 X as long as wide.

Pronotum 1.8 X as wide as long, broadest before base, with lateral sides rounded, more convergent forward than backward, with anterior side moderately emarginate, anterior angles rounded. Lateral callus weakly convex along entire length; lateral impression broad, shallow along entire length, deepened in anterior ½ and just before base. Pronotal disc evenly covered with dense, fine (0.02 mm) punctures. Lateral impression with 30 large (0.05–0.06 mm) punctures, partly coalescent basally. Eight moderate (0.03 mm) punctures present along outer margin of lateral callus. Pronotum entirely marginate and ciliate anteriorly, entirely marginate posteriorly. Anterior and posterior setiferous pores absent.

Prothoracic hypomeron weakly convex, with distinct impression and irregular wrinkles laterally. Anterior cavity and basal fold of prothoracic hypomeron absent.

Metasternum entirely marginate anteriorly.

Elytron without humeral callus. Elytral punctures moderate, (0.03 mm), arranged in paired (2–3, 4–5, 6–7, 8–9) distinct, regular rows, with intervals hardly convex, almost flat, covered by sparse fine (0.02 mm) punctures and obsolete wrinkles. 5^th row with 22 punctures. Scutellar row present. Sutural furrow distinct at apical slope. Elytral epipleuron inclined outside, visible along entire length in lateral view, sparsely ciliate apically.

Hind wing absent.

1^st–3^rd tarsomeres very broad, with entire sole. 1^st fore-tarsomere 1.1 X as wide as long, 0.9 X as wide as 3^rd fore-tarsomere, 3.1 X as wide as claw tarsomere.

Pygidium with very shallow longitudinal impression in basal ½ only.

Last abdominal ventrite simple, convex, with apical margin broadly shallowly emarginate, with furrow along apical margin.

Aedeagus (Figs 7E, 15F) tube-shaped, with dorsal side concave at most of its length, 5.3 X as long as wide, with parallel lateral sides along most of its length, slightly convergent at sides of apical orifice. Apical part slightly asymmetrical. Aedeagus without apical denticles on ventral side. Aedeagus moderately evenly curved in lateral view. Flagellum narrow, simple, exposed. Apical part with hoof-like thickening at the top dorsally. Apical orifice 0.4 X as long as aedeagus length, bearing sclerotized spoon-shaped appendage. Pores present laterally at level of apical orifice.

Body length 7.4 mm; aedeagus 3.2 mm long, 0.6 mm wide.

Paratype. Body length 7.4 mm.

Differential diagnosis. The new species is externally very similar with Ch. riccii sp. nov., and differs in aedeagus with apical part asymmetrical, bearing hoof-like thickening at the top and spoon-shaped appendage in the apical orifice.

Comments. Female is unknown.

Material examined: Holotype: China, N Sichuan Province, mts. 13 km NNE of Qinglang, alpine meadows, screes, 4,200 m, 32.20 N / 103.13 E, 24–27.7.2005, J. Kaláb leg.: male (ZIN); paratype: with the same label: 1 male (DC).

Description. Male (holotype) (Fig. 20C). Body oval, 1.6 X as long as wide. Body dark bronze, legs with greenish metallic reflection, antennae piceous with 1^st–2^nd antennomeres rufous below. Dorsum moderately shining, microreticulate and micropunctulate.

Supraorbital line long, not reaching antennal insertion. Base of antenna 2 X closer to clypeus than to eye. Last maxillary palpomere narrow, barrel-shaped, obliquely truncate apically, 1.3 X as long as wide, 1.1 X as long as penultimate palpomere, 1.1 X as wide as the latter. Antenna 0.4 X as long as body. 7^th antennomere 1.1 X as long as wide, 9^th antennomere 1.3 X as long as wide.

Pronotum 2.0 X as wide as long, broadest before base, with lateral sides rounded, more convergent forward than backward, with anterior side moderately emarginate, anterior angles rounded. Lateral callus weakly convex along entire length; lateral impression broad, shallow along entire length. Pronotal disc evenly covered with dense, fine to moderate (0.02–0.03 mm) punctures. Lateral impression with 25 large (0.05–0.06 mm) punctures. Eight moderate (0.03 mm) punctures present along outer margin of lateral callus. Pronotum entirely marginate and ciliate anteriorly, entirely marginate posteriorly. Anterior and posterior setiferous pores of pronotum absent.

Prothoracic hypomeron convex, with distinct impression and transverse wrinkles laterally. Anterior cavity and basal fold of prothoracic hypomeron absent.

Metasternum entirely marginate anteriorly.

Elytron without humeral callus. Elytral punctures moderate (0.03 mm), arranged in paired (2–3, 4–5, 6–7, 8–9) regular rows, with intervals slightly convex, covered by dense fine (0.02 mm) punctures and fine wrinkles. 5^th row with 33 punctures. Scutellar row reduced, with 2 punctures. Sutural furrow distinct at apical slope. Elytral epipleuron inclined outside, visible along entire length in lateral view, sparsely ciliate apically.

Hind wing absent.

1^st–3^rd tarsomeres very broad, with entire sole. 1^st fore-tarsomere 1.2 X as wide as long, as wide as 3^rd tarsomere, 3.6 X as wide as claw tarsomere.

Pygidium with very shallow longitudinal impression in basal ½ only.

Last abdominal ventrite simple, convex, with apical margin broadly emarginate, with furrow along apical margin.

Aedeagus (Figs 7D, 15I) tube-shaped, with dorsal side slightly concave at most of its length, 5.2 X as long as wide, with almost parallel lateral sides, hardly broadened toward apical orifice, then slightly constricted laterally at apical orifice, and finally elongate triangular. Aedeagus with 2 small denticles on ventral side before apex. Aedeagus moderately evenly curved in lateral view, then recurved upward at apex. Flagellum narrow, simple, exposed. Apical orifice 0.3 X as long as aedeagus length, with cover plate 0.2 X as long as apical orifice. Pores present near apex dorsally and laterally.

Body length 7.3 mm; aedeagus 3.1 mm long, 0.6 mm wide.

Paratype. Body length 7.3 mm.

Differential diagnosis. The new species is externally similar with Ch. luobinwangi, Ch. selene sp. nov., Ch. circe sp. nov., and especially, Ch. poloi sp. nov., and differs in aedeagus with long apical triangle, constricted at base, and strongly S-shaped in lateral view.

Comments. Female is unknown.

Material examined: Holotype: China, C Sichuan Province, pass 20 km WNW Fubian, alpine meadows, screes, 4,500 m, 31°20′ N / 102°16′ E, 30.7–2.8.2010, J. Kaláb leg.: male (ZIN); paratypes: with the same label: 1 female (BC); China, C Sichuan Province, pass 20 km WNW Fubian, alpine meadows, screes, 4,500 m, 31°20′ N / 102°16′ E, 8–11.8.2007, J. Kaláb leg.: 1 male (BC).

Description. Male (holotype) (Fig. 20A). Body elongate oval, 1.7 X as long as wide. Dorsal side dark bronze; underside and legs black with bronze reflection; antennae black with 1^st and 2^nd antennomeres rufous below apically, base of trochanters reddish. Dorsum shining, obsoletely microreticulate and micropunctulate.

Supraorbital line distinct, long, but not reaching antennal insertion. Base of antenna 2.7 X closer to clypeus than to eye. Last maxillary palpomere narrow, barrel-shaped, obliquely truncate at apex, 1.5 X as long as wide, 1.2 X as long as penultimate palpomere and 1.1 X as wide as the latter. Antenna 0.5 X as long as body. 7^th antennomere 1.4 X as long as wide, 9^th antennomere 1.2 X as long as wide.

Pronotum 1.9 X as wide as long, broadest basally, with lateral sides rounded, convergent forward, with anterior side weakly emarginate, anterior angles slightly rounded. Lateral callus weakly convex along entire length; lateral impression broad and shallow along most of its length, deepened in basal ⅓. Pronotal disc evenly covered with dense, fine (0.02 mm) punctures. Lateral impression with several moderate (0.03 mm) punctures near base. Several moderate (0.03 mm) punctures present also along outer margin of lateral callus. Pronotum entirely marginate and ciliate anteriorly, entirely marginate posteriorly. Anterior and posterior setiferous pores of pronotum absent.

Prothoracic hypomeron convex, without lateral impression, with obsolete wrinkles laterally. Anterior cavity of prothoracic hypomeron and basal fold absent.

Metasternum entirely marginate anteriorly.

Elytron without humeral callus. Elytral punctures fine to moderate (0.02–0.03 mm), partly arranged in irregular, some undulate, paired (2–3, 4–5, 6–7, 8–9) rows, with intervals flat. Broad intervals between pairs of rows covered with numerous fine (0.02 mm) punctures. Intervals with numerous obsolete wrinkles. 5^th row with 30 punctures. Scutellar row with 3 punctures. Sutural furrow distinct at apical slope. Elytral epipleuron inclined outside, visible along entire length in lateral view, with only few setae apically.

Hind wing absent.

1^st tarsomere moderately broadened in all tarsi, 2^nd and 3^rd narrow, 1^st–3^rd with entire sole. 1^st fore-tarsomere as wide as long, as wide as 3^rd tarsomere, 3.6 X as wide as claw tarsomere.

Pygidium with shallow, obsolete longitudinal impression in basal ½ only.

Last abdominal ventrite simple, convex, with apical margin broadly truncate and bearing furrow along apical margin.

Aedeagus (Fig. 4C, 15G) tube-shaped, with dorsal side concave along most of its length, 4 X as long as wide, with parallel sides along entire length, almost rounded at apex and bearing large, obtuse apical denticles ventrally. Aedeagus moderately curved in basal ½ and almost straight in apical ½ (in lateral view). Flagellum simple, narrow, exposed. Apical orifice 0.3 X as long as aedeagus length, cover plate 0.4 X as long as apical orifice. Pores present laterally at level of apical orifice.

Body length 6.0 mm; aedeagus 2.8 mm long, 0.7 mm wide.

Description of female (paratype). Body 6.0 mm long; 1^st–3^rd tarsomeres narrow, with entire sole; spermatheca absent.

Variability. Male (paratype) 6.4 mm long.

Differential diagnosis. The new species is externally close to Ch. hongyuanensis and differs in prothoracis hypomeron convex, and in aedeagus apically rounded (viewed dorsally), with ventral denticles, not anchor-shaped.

Comments. Female is ovoviviparous. Embryos with developed ocelli, mandibulae, setae on body segments, and legs were found inside the abdomen.

Material examined: Holotype: China, NW Sichuan Province, Chola Shan, 1.3 km NW of Chola Pass, 4,825 m, 31°57′00″ N / 98°54′57″ E, 18.7.2016, I. Belousov, I. Kabak leg.: male (ZIN); paratypes: with the same label: 1 male, 1 female (ZIN), 1 male, 1 female (BC), 3 males, 1 female (RC); China, NW Sichuan Province, Chola Shan, 8.1 km ENE of Chola Peak, 4,915 m, 31°48′18″ N / 99°9′16″ E, 17.7.2016, I. Belousov, I. Kabak leg.: 1 female (ZIN), 2 females (BC); China, Sichuan Province, NW Chola Pass, between Maniganggo et Dege, 29–31.7.2004, S. Murzin leg.: 1 male (BC).

Description. Male (holotype) (Fig. 16D). Body oval, 1.6 X as long as wide. Dorsal and ventral sides dark bronze; legs black; antennae piceous with 1^st antennomere below rufous, 2^nd and 3^rd mostly rufous. Head and pronotum sericeous, elytra moderately shining, dorsum micropunctulate and microreticulate.

Supraorbital line distinct above eye. Base of antenna 2.0 X closer to clypeus than to eye. Last maxillary palpomere narrow, barrel-shaped, 1.2 X as long as wide, similar with penultimate palpomere in length and width. Antenna 0.4 X as long as body. 7^th antennomere 1.1 X as long as wide, 9^th antennomere as long as wide.

Pronotum 2.0 X as wide as long, broadest at base, with lateral sides convergent forward, almost rectilinear in basal ½ and rounded in anterior ½, with anterior side moderately emarginate, anterior angles roundly triangular. Lateral callus weakly convex along entire length; lateral impression obsolete along entire length. Pronotum evenly covered with dense, fine (0.02 mm) punctures. Moderate or large punctures along outer margin of lateral callus absent. Pronotum entirely marginate and ciliate anteriorly, entirely marginate posteriorly. Anterior and posterior setiferous pores of pronotum absent.

Prothoracic hypomeron almost flat, without lateral impression, with obsolete wrinkles laterally. Anterior cavity of prothoracic hypomeron and basal fold absent.

Metasternum entirely marginate anteriorly.

Elytron with obsolete humeral callus. Elytral punctures moderate (0.03 mm), hollow-shaped, arranged in regular, equidistant, sparse rows. Intervals almost flat (obsoletely convex), with sparse fine (0.02 mm) punctures and sparse fine wrinkles. 5^th row with 15 punctures. Scutellar row with 1–4 punctures. Sutural furrow distinct in posterior part of apical slope. Elytral epipleuron inclined outside, visible along entire length in lateral view, with only few setae apically.

Hind wing absent.

1^st–3^rd tarsomeres slightly broadened, with entire sole. 1^st fore-tarsomere 0.9 X as wide as long, 0.9 X as wide as 3^rd tarsomere, 2.3 X as wide as claw tarsomere.

Pygidium with shallow, obsolete longitudinal impression in basal ½ only.

Last abdominal ventrite simple, convex, with apical margin shallowly emarginate and bearing furrow.

Aedeagus (Figs 7A, 15B) slightly flattened dorso-ventrally, 5 X as long as wide, with parallel lateral sides in basal ½ and slightly broadened in apical ½, with apical margin elongate triangular. Aedeagus with 2 small obtuse apical denticles on ventral side. Aedeagus arc-shaped, rather curved in basal ½ (viewed laterally), with flagellum narrow, simple, exposed. Apical orifice 0.3 X as long as aedeagus length, cover plate 0.1 X as long as apical orifice. Pores present at lateral sides at level of apical orifice (viewed laterally).

Body length 5.9 mm; aedeagus 2.5 mm long, 0.5 mm wide.

Description of female (paratypes). 1^st–3^rd tarsomeres narrow, with entire sole; last abdominal ventrite convex, slightly swollen, broadly truncate apically, elytral intervals sometimes slightly convex; spermatheca present.

Variability. Pronotal lateral sides evenly rounded in some specimens. 5^th elytral row with 12–17 punctures. Males 5.6 mm long. Females 7.0–7.1 mm long.

Differential diagnosis. The new species is externally close to Ch. pascolii sp. nov. and differs in aedeagus roundly triangular apically, pronotal lateral impression more shallow, obsolete, not deepened basally.

Material examined: Holotype: China, Yunnan Province, Tianbaoshan between Shangrila & Habaxue Shan, NE of Xiaozhongdian, 4,160 m, 27°35′47″ N / 99°54′34″ E, 21.5.2015, I. Belousov, I. Kabak, G. Davidian leg.: male (ZIN); paratypes: with the same label: 2 males (BC).

Description. Male (holotype) (Fig. 16A). Body elongate oval, 1.7 X as long as wide. Dorsal side dark bronze; underside and legs dark bronze with greenish metallic reflection; antennae piceous with 1^st–3^rd antennomeres rufous below. Dorsum moderately shining, microreticulate and micropunctulate.

Supraorbital line long, deeply impressed, not reaching antennal insertion. Base of antenna 1.7 X closer to clypeus than to eye. Last maxillary palpomere narrow, barrel-shaped, obliquely truncate apically, 1.5 X as long as wide, 1.5 X as long as penultimate palpomere, 1.1 X as wide as the latter. Antenna 0.5 X as long as body. 7^th antennomere 1.8 X as long as wide, 9^th antennomere 1.4 X as long as wide.

Pronotum 1.8 X as wide as long, broadest at mid-length, with lateral sides rounded, more convergent forward than backward, with anterior side moderately emarginate, anterior angles rounded. Lateral callus weakly convex along entire length; lateral impression broad and very weak along entire length. Pronotal disc evenly covered with dense, moderate (0.03 mm) punctures. Lateral impression with numerous (about 20) large (0.06–0.08 mm) punctures. Large punctures along outer margin of lateral callus absent. Pronotum entirely marginate and ciliate anteriorly, entirely marginate posteriorly. Anterior and posterior setiferous pores of pronotum absent.

Prothoracic hypomeron weakly impressed and covered with more or less transverse wrinkles laterally. Anterior cavity small, basal fold of prothoracic hypomeron absent.

Metasternum entirely marginate anteriorly.

Elytron without humeral callus. Elytral punctures large (0.05–0.06 mm), arranged in paired (2–3, 4–5, 6–7, 8–9) undulate and partly irregular rows. Intervals flat, covered by fine wrinkles. 5^th row with 25 punctures. Scutellar row with 5 punctures. Sutural furrow distinct at apical slope. Elytral epipleuron inclined outside, visible along entire length in lateral view, sparsely ciliate apically.

Hind wing absent.

1^st–3^rd tarsomeres very broad, with entire sole. 1^st fore-tarsomere 1.1 X as wide as long, as wide as 3^rd tarsomere, 3.2 X as wide as claw tarsomere.

Pygidium with weak, shallow longitudinal impression in basal ½ only.

Last abdominal ventrite simple, convex, with apical margin broadly truncate, with furrow along apical margin.

Aedeagus (Figs 3E, 8H) flattened, with dorsal side broadly concave along most of its length, 3.8 X as long as wide, with almost parallel lateral sides, hardly broadened toward apex, with apical margin shortly roundly triangular and bearing very short lamella. Aedeagus without apical denticles. Aedeagus moderately evenly curved in lateral view. Flagellum very narrow, simple, exposed. Apical orifice 0.3 X as long as aedeagus length, cover plate 0.1 X as long as apical orifice. Pores present near apex dorsally and ventrally.

Body length 6.4 mm; aedeagus 1.9 mm long, 0.5 mm wide.

Variability. Available males are rather uniform. One paratype has elytral puncture rows more irregular than those in the holotype. Body length 6.3–6.8 mm.

Differential diagnosis. The species is externally similar with Ch. luobinwangi and differs in the apical triangle of aedeagus shorter and bearing short apical lamella (apical triangle is longer and devoid of lamella in Ch. luobinwangi). Flagellum in the new species is narrower than in Ch. luobinwangi. Elytral puncture rows are paired in the new species; they are equidistant in Ch. luobinwangi.

Comments. Female is unknown.

The key is only suitable for identification of males. Females are known for 13 species of 20. A number of species are similar externally and differ only in the male characters such as the structure of the aedeagus, the shape of 1^st fore-tarsomere, and proportions of 7^th and 9^th antennomeres.

1. 1(4)Dorsal side purely black, without metallic reflection. Elytral punctures hollow-shaped. Male 1^st fore-trasomere narrow, 2.3 X as wide as claw tarsomere.
2. 2(3)Apical projection of aedeagus narrow, rounded (Figs 4B, 8D). 7^th antenomere 1.5 X as long as wide. Length 7.7–7.9 mm. Sichuan.

   • Ch. wangboi

3. 3(2)Apical projection of aedeagus trapeziform with projecting lateral corners (Fig. 4A). 7^th antenomere 0.9 X as long as wide. Length 7.0 mm. Sichuan.

   • Ch. boccaccioi

4. 4(1)Dorsal side dark metallic: bronze, violet, green, or coppery. Elytral punctures hollow-shaped or simple. Male 1^st fore-trasomere different: from narrow to very wide.
5. 5(14)Elytral punctures hollow-shaped.
6. 6(7)Lateral impression of pronotum without large punctures, which larger than those on disk. Male 1^st fore-trasomere narrow, 2.3 X as wide as claw tarsomere. Dorsum one-colored bronze. Elytral intervals almost flat. Aedeagus with apex elongate triangular, bearing 2 fine denticles on ventral side (Figs 7A, 15B). Length 5.9 mm. Sichuan.

   • Ch. romantsovi sp. nov.

7. 7(6)Lateral impression of pronotum with large punctures (0.05–0.09 mm), which larger than those on disk. Male 1^st fore-trasomere 2.4–4.5 X as wide as claw tarsomere. Dorsum bronze or green, violet, coppery, one-colored or two-colored. Elytral intervals from flat to convex.
8. 8(9)Male 1^st fore-trasomere narrower, 2.4 X as wide as claw tarsomere. Aedeagus with 2 fine apical denticles on ventral side; with apex rounded (Fig. 2A, B). Dorsum one-colored bronze. Elytral intervals almost flat. Length 4.7–5.2 mm. Sichuan.

   • Ch. patrikeevi sp. nov.

9. 9(8)Male 1^st fore-trasomere wider, 2.8–4.5 X as wide as claw tarsomere. Aedeagus without apical denticles. Dorsum metallic: green, violet, coppery, mostly two-colored, rarely (in Ch. nixiana) one-colored bronze. Elytral intervals convex.
10. 10(13)Lateral impression of pronotum with 25–30 large punctures. 5^th elytral row with 13–18 punctures. Male 1^st fore-trasomere 3.7–4.5 X as wide as claw tarsomere. Dorsum differently colored.
11. 11(12)9^th antennomere 1.7–1.9 X as long as wide. Aedeagus 5.2 X as long as wide, with apical projection roundly trapeziform, with apical orifice long, 0.6 X as long as aedeagus (Figs 1A–E, 8C). Dorsum entirely bronze or head and pronotum greenish bronze, and elytra coppery. Length 7.1–7.6 mm. Yunnan.

    • Ch. nixiana

12. 12(11)9^th antennomere 1.4 X as long as wide. Aedeagus 3.1 X as long as wide, with apex broadly truncate, with apical orifice short, 0.3 X as long as aedeagus (Figs 7B, 8E). Dorsum violet with green. Length 8.7 mm. Yunnan.

    • Ch. yunnana

13. 13(10)Lateral impression of pronotum with about 10 large punctures. 5^th elytral row with 10 punctures. Male 1^st fore-trasomere wide, 2.8 X as wide as claw tarsomere. Dorsum metallic green with violet. Aedeagus with apex broad, bearing 2 semi-transparent areas (Figs 4D, 8F). Length 7.4 mm. Yunnan.

    • Ch. aquamarina sp. nov.

14. 14(5)Elytral punctures simple (not hollow-shaped), with intervals flat or hardly convex.
15. 15(26)Aedeagus with apical denticles (fine or large, sometimes lobe-shaped) on ventral side. Dorsum bronze.
16. 16(21)Aedeagus with large apical denticles. These denticles sharp or obtuse (lobe-shaped).
17. 17(18)Pronotal lateral impression with about 35–40 moderate punctures along entire length. Flagellum of aedeagus broader, tube-shaped. 1^st fore-tarsomere 2.9 X as wide as claw tarsomere. 5^th elytral row with about 40 punctures. Aedeagus – Figs 7F, 8A. Length 6.6–7.1 mm. Sichuan.

    • Ch. hongyuanensis

18. 18(17)Pronotal lateral impression with less than 10 moderate or large punctures near base. Flagellum of aedeagus narrower, hair-shaped. 1^st fore-tarsomere 2.5–3.6 X as wide as claw tarsomere. 5^th elytral row with 16–30 punctures.
19. 19(20)7^th antennomere 1.0–1.1 X as long as wide. 1^st fore-tarsomere 2.5–2.9 X as wide as claw tarsomere. Elytral punctures surrounded by large (0.2 mm) spot of strong reticulation. 5^th elytral row with 16–22 punctures. Aedeagus with apical margin broadly truncate and bearing 2 lobe-shaped denticles on ventral side (Figs 1J, 15A). Length 6.1–6.8 mm. Sichuan.

    • Ch. margaritifera sp. nov.

20. 20(19)7^th antennomere 1.4 X as long as wide. 1^st fore-tarsomere 3.6 X as wide as claw tarsomere. Elytral punctures simple, not surrounded by large spot of strong reticulation. 5^th elytral row with 30 punctures. Aedeagus with apical margin roundly triangular and bearing 2 lobe-shaped denticles on ventral side (Figs 4C, 15G). Length 6.0–6.4 mm. Sichuan.

    • Ch. rillingi sp. nov.

21. 21(16)Aedeagus with fine apical denticles.
22. 22(23)9^th antennomere 1.1 X as long as wide. 1^st fore-tarsomere 2.6 X as wide as claw tarsomere. Aedeagus with apex elongate triangular, narrowly truncate (Figs 7C, 15A). Length 5.7–6.0 mm. Sichuan.

    • Ch. pascolii sp. nov.

23. 23(22)9^th antennomere 1.3–1.5 X as long as wide. 1^st fore-tarsomere 3.2–3.6 X as wide as claw tarsomere.
24. 24(25)7^th antennomere about as long as wide; 9^th antennomere 1.3 X as long as wide. Pronotal lateral impression with 25 large punctures. Elytral rows paired. 1^st fore-tarsomere 3.6 X as wide as claw tarsomere. Aedeagus with apex elongate triangular, narrowly truncate (Figs 7D, 15I). Length 7.3 mm. Sichuan.

    • Ch. riccii sp. nov.

25. 25(24)7^th antennomere 1.3 X as long as wide; 9^th antennomere 1.5 X as long as wide. Pronotal lateral impression with about 10 moderate punctures. Elytral rows equidistant. 1^st fore-tarsomere 3.2 X as wide as claw tarsomere. Aedeagus with apex truncate and bearing shallow emargination (Figs 1I, 8G). Length 6.0–6.8 mm. Yunnan.

    • Ch. circe sp. nov.

26. 26(15)Aedeagus without any apical denticles. Dorsum entirely bronze or coppery with green pronotum.
27. 27(32)7^th antennomere 1.0–1.1 X as long as wide. Flagellum of aedeagus absent or present, hair-shaped or tube-shaped. Dorsum bronze or coppery with pronotum green.
28. 28(29)Pronotal lateral impression without large punctures. Aedeagus without flagellum (Figs 3D, 15C). Dorsum bronze. Length 5.5–5.8 mm. Sichuan.

    • Ch. pakhomovae sp. nov.

29. 29(28)Pronotal lateral impression with large punctures. Aedeagus with flagellum. Dorsum bronze or coppery with pronotum green.
30. 30(31)Flagellum of aedeagus wider, tube-shaped (Figs 3F, 8I). Dorsum coppery with pronotum green. 1^st fore-tarsomere 2.5 X as wide as claw tarsomere. 5^th elytral row with 35 punctures. Length 6.8 mm. Yunnan.

    • Ch. leda sp. nov.

31. 31(30)Flagellum of aedeagus narrower, hair-shaped (Figs 4B, 8E). Dorsum one-colored bronze. 1^st fore-tarsomere 2.8 X as wide as claw tarsomere. 5^th elytral row with 16–21 punctures. Length 5.4–6.2 mm. Sichuan.

    • Ch. luobinwangi

32. 32(27)7^th antennomere 1.3–1.8 X as long as wide. Flagellum of aedeagus narrow, hair-shaped. Dorsum bronze.
33. 33(36)7^th antennomere 1.6–1.8 X as long as wide. 1^st fore-tarsomere 3.1–3.2 X as wide as claw tarsomere.
34. 34(35)7^th antennomere 1.6 X as long as wide; 9^th antennomere 1.7 X as long as wide. Elytral rows with moderate (0.03 mm) punctures. Aedeagus with hoof-like thickening at apex (Figs 7E, 15F). Length 7.4 mm. Yunnan.

    • Ch. poloi sp. nov.

35. 35(34)7^th antennomere 1.8 X as long as wide; 9^th antennomere 1.4 X as long as wide. Elytral rows with large (0.05–0.06 mm) punctures. Aedeagus with apex roundly triangular and bearing short lamella (Figs 3E, 8H). Length 6.4 mm. Yunnan.

    • Ch. selene sp. nov.

36. 36(33)7^th antennomere 1.3-1.4 X as long as wide. 1^st fore-tarsomere 1.9–2.3 X as wide as claw tarsomere.
37. 37(38)9^th antennomere 1.6 X as long as wide. Aedeagus longer, 4.0 X as long as wide (Figs 3A–C, 15 D). Length 5.6–8.4 mm. Yunnan.

    • Ch. konstantinovi sp. nov.

38. 38(37)9^th antennomere 1.2 X as long as wide. Aedeagus shorter, 2.7 X as long as wide (Fig 1K, 15E). Length 5.4–5.8 mm. Yunnan.

    • Ch. luluni sp. nov.